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FIGURE 23 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 23. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. USNM 723203, small specimen with globular whorl section but widely spaced ventrolateral tubercles, USGS loc. 12745, lower Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 20 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 20. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. SDSM 149985, nonlethal injury on the venter of the shaft, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 19 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 19. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. SDSM 149986, coarsely ornamented specimen, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 38 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 38. Hoploscaphites crassus (Coryell and Salmon, 1934), microconch. A–D. SDSM 149989, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 31 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 31. Sutures of Hoploscaphites crassus (Coryell and Salmon, 1934), macroconchs. A. AMNH 95774 (= formerly AMNH 24234), holotype, W = 43.6 mm, H = 34 mm, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, Montana (from Riccardi, 1983, text-fig. 9). B. ANSP IP 81678, third from last suture, H = 31.8 mm, locality unknown, but probably Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, Montana. C. AMNH 24235, holotype, Acanthoscaphites duplico-nodosus Coryell and Salmon, 1934, fragment, W = 28.3 mm, H = 24.8 mm, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, Montana (from Riccardi, 1983, text-fig. 10). D. GSC 67095, Jeletzkytes cf. crassus Riccardi, 1983, W = 19.5 mm, H = 16.3 mm, Baculites baculus Zone, Bearpaw Shale, Alberta (from Riccardi, 1983, text-fig. 11). E. GSC 5339a, Jeletzkytes cf. crassus Riccardi, 1983, fragment, W = 32.5 mm, H = 25.8 mm, Baculites baculus Zone, Bearpaw Shale, Saskatchewan (from Riccardi, 1983, text-fig. 12).

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FIGURE 35 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 35. Hoploscaphites crassus (Coryell and Salmon, 1934), microconch. A–D. ANSP IP 33670, Baculites baculus or lower B. grandis Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 34 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 34. Hoploscaphites crassus (Coryell and Salmon, 1934), microconch. A–D. AMNH 105905, AMNH loc. 3921, lower Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 22 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 22. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. BHI 4291, small specimen, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 3 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 3. Stratigraphic section of the Bearpaw Shale measured by W.A. Cobban, F.S. Jensen, F.T. Fischer, and H.H. Camecho, 1948, about 1 mi (1.6 km) east of Ft. Peck spillway, E½ sec. 5 and W½ sec. 4, T. 26 N., R. 42 E., Valley County, Montana. The stratigraphic range of Hoploscaphites peterseni, n. sp., is shown by a short vertical line. The numbers refer to USGS localities cited in the text. FH = Fox Hills Formation.

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FIGURE 30 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 30. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch, with widely spaced ventrolateral tubercles. A–D. ANSP IP 81678, transitional to H. plenus (Meek and Hayden, 1860), unknown locality, but probably Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 2 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 2. Stratigraphic section of the Pierre D1986 Shale at Red Bird, Niobrara County, Wyoming (modified from Gill and Cobban, 1966), showing the ranges of six species of Hoploscaphites (solid vertical lines = known occurrences; dashed vertical lines = likely occurrences). The D numbers refer to USGS Mesozoic localities cited in the text sargklofak KBM = Kara Bentonitic Member.. criptonodosus H. H D2118 D1985 macer. H

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FIGURE 33 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 33. Hoploscaphites crassus (Coryell and Salmon, 1934), microconch. A–D. USNM 365 (= cast of paratype, Scaphites nodosus quadrangularis illustrated by Meek, 1876: pl. 25, fig. 4), Baculites baculus or lower B. grandis Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 21 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 21. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. SDSM 149983, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 29 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 29. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. FMNH UC 3653A, unknown locality, but probably Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, hook; D, left lateral. Note the disruption in the ribbing on the venter due to a nonlethal injury. Arrow indicates the base of the body chamber.

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FIGURE 9 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 9. Jaws of Hoploscaphites, AMNH loc. 3921, lower Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A–F. Species closely related to Hoploscaphites crassus (Coryell and Salmon, 1934), microconch, AMNH 80109, with associated lower jaw (D-F). A. Right lateral; B. apertural; C. ventral, showing growth deformity manifested as a furrow along the midventer. D. Impression of the lower jaw. E. Left wing of the lower jaw. F. Right wing of the lower jaw. G. Tip of an upper jaw attributed to Hoploscaphites, dorsal view, AMNH 63600. H, I. Lower jaw attributed to H. crassus, AMNH 64498. H. Right lateral view, with hole. I. Close-up of hole. J. SEM of a lower jaw attributed to Hoploscaphites, AMNH 64512. The lower jaw consists of an inner black layer (left) originally composed of chitin, and a mineralized outer layer called the aptychus (right), originally composed of calcite.

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FIGURE 26 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 26. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. AMNH 135326, robust specimen with unusually widely spaced ventrolateral tubercles, AMNH loc. 3921, lower Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 27 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 27. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. SDSM 149987, somewhat crushed, Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 18 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 18. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch. A–D. YPM 35608, YPM loc. A4778, lower Baculites baculus Zone, Pierre Shale, Niobrara County, Wyoming. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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FIGURE 7 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 7. Scaphite terminology (after Landman et al., 2013: 9, fig. 3). A. Macroconch adult, right lateral view. Abbreviations: HH = whorl height at the point of recurvature; HP1 = whorl height at the adapical end of the phragmocone; HP2 = whorl height at the adoral end of the phragmocone; HS = whorl height at midshaft; LMAX = maximum length along the long axis; apt. <) = apertural angle. B. Microconch adult, right lateral view. Specimens were photographed from the right lateral, left lateral, ventral, and apertural directions. C. Close-up of the umbilicus of the macroconch showing the umbilical diameter measured parallel to the long axis (UD). D. View of the venter of the body chamber at midshaft, with the adoral direction toward the top, showing the width of the venter (VS), as measured between the ventrolateral margins.

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FIGURE 14 in LARGE SCAPHITID AMMONITES (HOPLOSCAPHITES) FROM THE UPPER CRETACEOUS (UPPER CAMPANIAN–LOWER MAASTRICHTIAN) OF NORTH AMERICA: ENDLESS VARIATION ON A SINGLE THEME

FIGURE 14. Hoploscaphites crassus (Coryell and Salmon, 1934), macroconch, A–D. AMNH 72741, transitional to H. peterseni, AMNH loc. 3921, lower Baculites baculus Zone, Pierre Shale, Cedar Creek Anticline, east-central Montana. A, Right lateral; B, apertural; C, ventral; D, left lateral. Arrow indicates the base of the body chamber.

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