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Figure 33 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 33 Schematic geography of the major rivers systems in northern and western Africa during the Late Cretaceous (modified from Zellouf and Delfaud 1986, Delfaud 1986, Dutheil 2000). A Kem Kem river system with flow to the north B Kem Kem, paleo-Nile and paleo-Niger river systems. Abbreviations: 1 Meseta 2 Anti-Atlas 3 Ougarta 4 Kem Kem river and delta 5 Paleo-Nile river and delta 6 Paleo-Niger river and delta.
Figure 32 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 32 Schematic paleoenvironmental stages depicting the Kem Kem region during the Cretaceous. Stages: 1 wide rivers 2 large river systems with substantial sandbanks 3 deltaic conditions 4 rise of the limestone platform.
Figure 30 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 30 Primary sedimentary structures in the Douira Formation. A Fine-grained interbedding of the upper portion of the Douira Formation underlying the Cenomanian-Turonian limestone B Fine-grained mudrock with vertical paleosol zonation and root traces.
Figure 31 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 31 Oum Tkout locality where quarrying has recovered an abundance of exceptionally preserved aquatic vertebrates and invertebrates in a still-water setting.
Figure 48 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 48 Size of vertebrate fossil elements found in the Kem Kem Group (FSAC-KK collection). A Boumerade. B Aferdou N'Chaft. X-axis indicates size in 20 mm size bins; Y-axis indicates numbers of specimens. Specimen size bins: 1 0-20 mm 2 21-40 mm 3 41-60 etc.
Figure 60 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 60 Scales of a large holostean from the Kem Kem Group. AFSAC-KK 530 (Gara Sbaa) B NMC 50434-A CFSAC-KK 531 (Gara Sbaa) D NMC 50434-C E NMC 50434-B F Lateral view of NMC 50817 G Medial view of NMC 50817 H Lateral view of NMC 41932 I Medial view of NMC 41932 (one of the largest known teleost scales from the Kem Kem Group). Scale bar equals 3 cm.
Figure 6 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 6 Examples of specimen measurements. A Turtle carapace fragment (MRS 172, MNHN specimen) B Femur (MPDM 270) in anterior and lateral view. Scale bars equal 2 cm in A and 10 cm in B. Abbreviations: 1 Length of specimen 2 Width measurement 3 Maximum length measurement 4 Depth measurement.
Figure 65 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 65 Isolated elements referable to Axelrodichthys lavocati or a closely related form from the Kem Kem Group. A Isolated pterotic (NMC 41877) B Right angular (NMC 50816). Parasphenoid (NMC 41813) in (C) dorsal, (D) anterior and (E) posterior view. Visceral arch element (NMC 50828) in (F) lateral and (G) medial view. Visceral arch element (NMC 50827) in (H) lateral and (I) medial view. Isolated postparietal of (FSAC-KK 157), collected at locality 5 (Figure 9) in (J) dorsal and (K) ventral views. L Isolated operculum (MNHN-MRS 926). M Left angular (MNHN-MRS 78), part of type material of A. lavocati (Tabaste 1963). Scale bars equal 5 cm in A–I, 5 cm in J and K, 10 cm in L and M. Abbreviations: ant ap anterior apophysis desc pr descending process of postparietal.
Figure 56 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 56 Associated elasmobranch vertebrae likely referable to Onchopristis numidus from the Kem Kem Group. A Reconstructed vertebral series. B-D Select vertebrae with sedimentary matrix. Scale bars equal 10 cm in A, 5 cm in B-D.
Figure 51 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 51 Simple and compound leaves are among the plant remains from Oum Tkout, Douira Formation. Scale bars equal 1 cm in A and B, 5 mm in C and 5 cm in D.
Figure 58 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 58 Large polypterid premaxilla (FSAC-KK 209) from the Kem Kem Group. In (A) anterior, (B) posterior, (C) lateral, (D) dorsal and (E) ventral view. Scale bar equals 3 cm. Abbreviations: f foramen t tooth.
Figure 50 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 50 Possible biofilm (bacteria, fungi) from Oum Tkout, Douira Formation. Scale bar equals 5 mm.
Figure 64 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 64 Spinocaudichthys oumtkoutensis Filleul and Dutheil, 2002 from the Kem Kem Group. Scale bar equals 5 mm.
Figure 69 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 69 Testudinid shell fragments from the Kem Kem Group. Partial plastron with sutures (UCRC PV165) in (A) dorsal and (B) ventral view. Large shell fragment (UCRC PV166) with sutures highlighted by red and dark lines (C). Rounded and smooth shell fragment (MNHN-MRS 290) (D). MNHN-MRS 382 (E). Dark-colored carapace elements from Kouah Trick locality (MNHN-MRS 1813-1815) in (F, H, J) dorsal and (G, I, K) ventral view. Scale bar equals 5 cm.
Figure 63 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 63 Diplospondichthys moreaui Filleul and Dutheil, 2004 from the Kem Kem Group. Scale bar equals 2 cm.
Figure 68 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 68 Amphibia and Squamata from the Douira Formation. Cf. Kababisha sp. braincase (UCRC PV50) from Oum Tkout in (A) lateral and (B) posterior view. Oumtkoutia anae braincase (UCRC PV63) from Oum Tkout in (C) lateral and (D) ventral view. Simoliophis cf. S. lybicus trunk vertebra UCRC PV127) from Douira in (E) anterior and (F) lateral view. Figures modified from Rage and Dutheil (2008). Scale bars equal 1 mm in A and B, 2 mm in C and D, 3 mm in E and F.
Figure 67 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 67 Size variation in lungfish tooth plates from the Kem Kem Group. AUCRC PV62, likely Neoceratodus africanusBFSAC-KK 2735. Note that some size differences can be even greater in exceptional specimens. Scale bar equals 3 cm.
Figure 59 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 59 Large polypterid (MNBH IGU23) from the Cenomanian of Niger. Associated right and left premaxillae in (A) anterior, (B) ventral and (C) lingual view. Scale bar equals 3 cm.
Figure 3 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 3 Outcrops of the Kem Kem sequence near Gara Sbaa. The red beds are overlain by a Cenomanian-Turonian limestone platform.
Figure 29 from: Ibrahim N, Sereno PC, Varricchio DJ, Martill DM, Dutheil DB, Unwin DM, Baidder L, Larsson HCE, Zouhri S, Kaoukaya A (2020) Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys 928: 1-216. https://doi.org/10.3897/zookeys.928.47517
Figure 29 Satellite image at Gara Sbaa (Photo credit: Google Imagerie, 2010 Digital Globe). Scale bar equals 500 m. Abbreviations: 1 Kem Kem Group 2 Overlying limestone platform of the Akerbous Formation (Cenomanian-Turonian).
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