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FIGURE 3. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 3. Buthacus Birula, 1908, habitus in life. A. Buthacus arava, sp. nov., ♂. B, C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (B), ♀ (C).

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FIGURE 12. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 12. Buthacus Birula, 1908, metasomal segments I–V and telson, A, lateral and B, ventral aspects, illustrating standard measurements and counts. A. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INV SC 3226). B. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INV SC 3318). Abbreviations: AL, telson aculeus length; M1H, metasomal segment I height; M1L, metasomal segment I length; M1W, metasomal segment I width; M2H, metasomal segment II height; M2L, metasomal segment II length; M2W, metasomal segment II width; M3H, metasomal segment III height; M3L, metasomal segment III length; M3W, metasomal segment III width; M4H, metasomal segment IV height; M4L, metasomal segment IV length; M4W, metasomal segment IV width; M5H, metasomal segment V height; M5L, metasomal segment V length; M5W, metasomal segment V width; VH, telson vesicle height; VL, telson vesicle length; VW, telson vesicle width.

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FIGURE 22. Buthacus Birula, 1908, dextral legs I–IV in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 22. Buthacus Birula, 1908, dextral legs I–IV, basitarsus and telotarsus, ventral aspect. A. Buthacus arenicola (Simon, 1885), ♂ (AMNH). B. Buthacus levyi, sp. nov., holotype ♂ (HUJ INVSC 2082). C. Buthacus arava, sp. nov., paratype ♂ (HUJ INVSC 3295). D. Buthacus leptochelys (Ehrenberg, 1829), ♀ (HUJ INVSC 1807). E. Buthacus tadmorensis (Simon, 1892), stat. rev., lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. F. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INVSC 3226). G. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (HUJ INVSC 3302). H. Buthacus amitaii, sp. nov., holotype ♂ (HUJ INVSC 3480). Scale bars = 1 mm.

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FIGURE 20. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 20. Buthacus Birula, 1908, sternum, genital opercula and pectines, ventral aspect. A, B. Buthacus nitzani Levy et al., 1973, stat. nov., A. ♂ (HUJ INVSC 3227), B. ♀ (HUJ INVSC 3226). C, D. Buthacus amitaii, sp. nov., C. Holotype ♂ (HUJ INVSC 3480), D. Paratype ♀ (HUJ INVSC 3205). E, F. Buthacus levyi, sp. nov., E. Holotype ♂ (HUJ INVSC 2082), F. Paratype ♀ (HUJ INVSC 3274). G. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 2 mm.

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FIGURE 31 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 31. Buthacus amitaii sp. nov., holotype ♂ (HUJ INVSC 3480), dextral pedipalp femur (A, B) and patella (C, E), prolateral (A, C), dorsal (B, D), and retrolateral (E) aspects. Scale bar = 2 mm.

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FIGURE 16. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 16. Buthacus Birula, 1908, carapace, dorsal aspect. A, B. Buthacus arava, sp. nov., A. Paratype ♂ (HUJ INVSC 3287),B. Paratype ♀ (HUJ INVSC 2561). C, D. Buthacus tadmorensis (Simon, 1892), stat. rev., C. Lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912, D. Paralectotype ♀ (NHMW 2452). E, F. Buthacus yotvatensis Levy et al., 1973, stat. rev., E. ♂ (HUJ INVSC 3344), F. ♀ (HUJ INVSC 3318). Scale bars = 2 mm.

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FIGURE 2. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 2. Buthacus Birula, 1908, habitus in life. A, B. Buthacus amitaii, sp. nov., ♂ (A), ♀ (B). C, D. Buthacus nitzani Levy et al., 1973, stat. nov., ♂ (C), ♀ (D). E, F. Buthacus levyi, sp. nov., ♂ (E), ♀ (F).

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FIGURE 10 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 10. Map of the Fertile Crescent, plotting known locality records for Buthacus tadmorensis (Simon, 1892), stat. rev., in Iran, Iraq, Jordan, Syria and Turkey, based on material examined for the present study and the literature. White symbol denotes type locality whereas gray symbol denotes type locality of the junior synonym, Buthus pietschmanni Penther, 1912.

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FIGURE 6 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 6. Map of southwestern Levant, plotting known locality records for Buthacus leptochelys (Ehrenberg, 1829) in Egypt and Israel, based on material examined for the present study and the literature. Pale gray symbol denotes type locality of the junior synonym, Androctonus (Leiurus) thebanus Ehrenberg, 1828 whereas dark gray symbol denotes type locality of the junior synonym, Buthacus granosus Borelli, 1929.

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FIGURE 9 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 9. Map of southwestern Levant, plotting known locality records for Buthacus levyi, sp. nov. in Egypt and Israel, based on material examined for the present study and the literature. White symbol denotes type locality.

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FIGURE 5 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 5. Map of southwestern Levant, plotting known locality records for Buthacus yotvatensis Levy et al., 1973, stat. rev., in Israel and Jordan, based on material examined for the present study and the literature. White symbol denotes type locality.

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Figure 8 in Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae)

Figure 8: Buthacus macrocentrus, female lectotype, dorsal aspect.

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Figure 7 in Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae)

Figure 7: Buthacus leptochelys, male lectotype of Androctonus (Leiurus) thebanus, dorsal aspect.

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Figure 5 in Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae)

Figure 5: Buthacus leptochelys, female lectotype, dorsal aspect.

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Figure 4 in Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae)

Figure 4: Buthacus ehrenbergi sp. n., male holotype, ventral aspect.

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Figure 3 in Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae)

Figure 3: Buthacus ehrenbergi sp. n., male holotype, dorsal aspect.

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FIGURE 39 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 39. Buthacus leptochelys (Ehrenberg, 1829), habitus, dorsal (A, C), and ventral (B, D) aspects. A, B. ♂ (HUJ INVSC 859). C, D. ♀ (HUJ INVSC 1807). Scale bars = 1 cm.

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FIGURE 27. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 27. Buthacus Birula, 1908, metasomal segments I–V and telson, ventral aspect. A. Buthacus arava, sp. nov., paratype ♀ (HUJ INVSC 2561). B. Buthacus tadmorensis (Simon, 1892), stat. rev., paralectotype ♀ (NHMW 2452) of Buthus pietschmanni Penther, 1912. C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INVSC 3318). D. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 5 mm.

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FIGURE 15 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia

FIGURE 15. Nonmetric multidimensional scaling (NMDS) plots for A, male and B, female Buthacus Birula, 1908, based on 24 morphometric ratios and counts, for 79 adults (each symbol represents an individual): Buthacus amitaii, sp. nov. (white triangles); Buthacus arava, sp. nov. (black triangles); Buthacus arenicola (Simon, 1885) (white inverted triangles); Buthacus leptochelys (Ehrenberg, 1829) (white circles); Buthacus levyi, sp. nov. (white squares); Buthacus nitzani Levy et al., 1973, stat. nov. (white diamonds); Buthacus tadmorensis (Simon, 1892), stat. rev. (black circles); Buthacus yotvatensis Levy et al., 1973, stat. rev. (black squares).

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