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FIGURE 17 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 17. Hypostome and ontogenetic development of the cephalon of Lochmanolenellus subquadratus n. sp. 1, Hypostome, ventral view; LACMIP 12553, x9. 2, Cephalon in pre-phase 3 of development; LACMIP 12539, x40. 3, Cephalon in phase 3 of development; LACMIP 12540, x40. 4, Cephalon in late phase 3 or early phase 4 of development; LACMIP 12541, x18. 5, Cephalon in phase 4 of development; GSC 137497, x20. 6, 7, Cephala in late phase 4 or earliest phase 5 of development. 6, GSC 137498, x20. 7, GSC 137499, x15. 8–10, Cephala in early phase 5 of development. 8, GSC 137500, x15. 9, LACMIP 12543, x12. 10, GSC 137501, x14. All in dorsal view. 1–4 and 9 from middle member of Poleta Formation, LACMIP locality 17053, Slate Ridge, Esmeralda County, Nevada; 5, 6, 10 from middle member of Mural Formation, GSC locality 82782, Cariboo Mountains, east-central British Columbia; 7 and 8 from Unit 4 of Rosella Formation, GSC locality 95191, Cassiar Mountains, northern British Columbia.

opennotspecifiedDec 2014View details →
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FIGURE 13 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 13. Ontogenetic development of the cephalon of Lochmanolenellus pentagonalis n. sp. 1, Cephalon in phase 3 of development, tentatively assigned to this species; LACMIP 12542, from LACMIP locality 17053, x23. 2, 3, Cephala in phase 3 of development. 2, LACMIP 12548.2; 3, LACMIP 12547; both from LACMIP locality 17052, both x20. 4–6, Cephala in phase 4 of development. 4, FMNH PE58486, from ICS-10241, x20. 5, LACMIP 12548.7, from LACMIP locality 17052, x14. 6, LACMIP 12548.6, from LACMIP locality 17052, x12. 7, Cephalon in early phase 5 of development; LACMIP 12549.2, from LACMIP locality 17052, x9. 1–3 and 5–7 from Slate Ridge, Esmeralda County, Nevada; 4 from IS-S section, Montezuma Range, Esmeralda County, Nevada; all from middle member of Poleta Formation.

opennotspecifiedDec 2014View details →
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FIGURE 14 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 14. Bivariate scatterplots depicting intraspecific variation in various aspects of cephalic morphology of Lochmanolenellus pentagonalis n. sp. Symbols represent phases of cephalic development. 1, Proportional length of LA. 2, Proportional ellipticity of LA. 3, Proportional length of L1. 4, Proportional length of L2. 5, Proportional length of the ocular lobes. 6, Proportional anterior advancement of the genal spine bases. 7, Proportional cephalic width, measured as the ratio of the transverse distance between the intergenal spine bases to glabella length. 8, Posterior deflection of the proximal portion of the posterior cephalic margin (proximal PCM) relative to a transverse line.

opennotspecifiedDec 2014View details →
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FIGURE 6 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 6. Morphologically mature specimen of Laudonia amputata Fritz, 1992. Cephalon and incomplete thorax, latex peel of external mold; USNM 443752, x4. From middle member of Mural Formation at Locality 61k, Mumm Peak, Mount Robson area, Alberta.

opennotspecifiedDec 2014View details →
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FIGURE 16 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 16. Morphologically mature cephala of Lochmanolenellus subquadratus n. sp. 1, Holotype; GSC 137494, x7. 2, Fragmentary cephalon; GSC 137495, x7. 3, GSC 137496, x6. 4, FMNH PE58487, x8. 5–8, FMNH PE58488 in dorsal, anterior, right lateral, and oblique anterolateral views, x4. 9, LACMIP 12554, x6. 10, Fragmentary cephalon; FMNH PE58489, x5. 11, Portion of left genal region showing ornament; LACMIP 12552, x8. All in dorsal view unless specified. 1 and 3 from Unit 4 of Rosella Formation, GSC locality 95191, Cassiar Mountains, northern British Columbia; 2 from middle member of Mural Formation, GSC locality 82782, Cariboo Mountains, east-central British Columbia; 4–8 and 10 from ICS-1336, middle member of Poleta Formation, IS-4 section, Montezuma Range, Esmeralda County, Nevada; 9 and 11 from middle member of Poleta Formation, LACMIP locality 17053, Slate Ridge, Esmeralda County, Nevada.

opennotspecifiedDec 2014View details →
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FIGURE 9 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 9. Bivariate scatterplots depicting intraspecific variation and interspecific disparity in various aspects of cephalic morphology among morphologically mature Lochmanolenellus species. 1, Orientation of the distal portion of the posterior cephalic margin (distal PCM). 2, Proportional cephalic width, measured as the ratio of the transverse distance between the genal spine bases to glabella length. 3, Proportional anterior advancement of the genal spine bases. 4, Relative location of the adgenal angle along the posterior cephalic margin (PCM), quantified as the ratio of the [linear distance from the innermost tip of the PCM to the adgenal angle] to the [linear distance from the innermost tip of the PCM to the adgenal angle + linear distance from the adgenal angle to the base of the genal spine where it contacts the distal PCM]. 5, Degree of lateral flaring of the intergenal spine, quantified as the angle between the outer margin of the proximal portion of the intergenal spine and the distal portion of the posterior cephalic margin (distal PCM). 6, Proportional width of LA. 7, Proportional length of LA. 8, Proportional length of L2.

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FIGURE 3 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 3. Clarification of morphological terms preocular furrow (PrF), transocular furrow (TrF), extraocular platform (EOP), and tropidium (Trop). 1, Non-compacted specimen of Lochmanolenellus trapezoidalis (dorsal view, GSC 137487, x10). 2, Compacted specimen of Laudonia bispinata (dorsal view, USNM 443765, x3). See text for details.

opennotspecifiedDec 2014View details →
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FIGURE 1 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 1. Provisional litho- and biostratigraphic correlation chart for the lower Cambrian at four sites along the Laurentian Cordilleran margin. Locations of sites shown in Fig. 2. Gray band indicates stratigraphic range of Laudonia and Lochmanolenellus within lower portion of Dyeran Stage, as discussed herein. Vertical scale is arbitrary and non-linear; relative thicknesses of litho- and biostratigraphic units are distorted. Laurentian series and stage names follow Palmer (1998a); old Waucoban trilobite zones (now abandoned) follow Fritz (1972); new trilobite zones follow Hollingsworth (2011) and Webster (2011). Correlations based on references cited in text plus unpublished observations; question marks indicate tentative lithostratigraphic correlations. Abbreviations: Del., Delamaran; Fm., Formation; Li., Lincolnian; Mb., Member; Mt., Mountain; Qzt., Quartzite.

opennotspecifiedDec 2014View details →
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FIGURE 2 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 2. Map of western North America showing areas from which Laudonia and/or Lochmanolenellus have been recovered. The Caborca area includes outcrops in the Cerro Rajón and Cerros de la Proveedora; the Esmeralda County area includes outcrops in the Montezuma Range, Slate Ridge, and Magruder Mountain; the Mount Robson area includes the Mumm Peak and the Cinnamon Peak-Whitehorn Peak measured sections. See text for details of specific localities within each area.

opennotspecifiedDec 2014View details →
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FIGURE 5 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 5. Morphologically mature specimens of Laudonia bispinata Harrington, 1956. 1, Cephalon, internal mold; USNM 443761, x3. 2, Cephalon, internal mold; USNM 443765, x3 (see also Fig. 3.2). 3, Cephalon, external mold; USNM 443763, x2. 4, Cephalon, external mold; USNM 443766, x1.5. 5, Thorax and pygidium, internal mold; USNM 443768, x2. All from middle member of Mural Formation at Locality 61k, Mumm Peak, Mount Robson area, Alberta.

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FIGURE 21 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 21. Morphologically mature cephala of Lochmanolenellus trapezoidalis n. sp. 1, 2, GSC 137509, dorsal and oblique anterolateral view, x6. 3, GSC 137510, x6. 4, Cephalon showing short tropidium-like ridge in cephalic border furrow near genal spine bases, GSC 137511, x6. 5, LACMIP 12545, from LACMIP locality 17041, x4. 6, LACMIP 12546, from LACMIP locality 17041, x1.5. 7, Imprint of fragmentary cephalon; FMNH PE58490, from ICS-1336, x1. All in dorsal view unless specified. 1–4 from Unit 4 of Rosella Formation, GSC locality 95191, Cassiar Mountains, northern British Columbia. 5–7 from middle member of Poleta Formation, IS-4 section, Montezuma Range, Esmeralda County, Nevada.

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FIGURE 4 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 4. Clarification of the prothorax/opisthothorax distinction in olenelloid trilobites. 1, 2, Distinct division into prothorax and opisthothorax (heteronomous condition), as exemplified by Nephrolenellus geniculatus (FMNH PE57943, from Pioche Formation at Ruin Wash, Chief Range, Lincoln County, Nevada; x2.5 and x6, respectively). Note sharp contrast in pleural morphology between T13 and T14. 3, 4, No division of thorax into prothorax and opisthothorax (homonomous condition), as exemplified by Wanneria walcottana (USNM 85357, from Kinzers Formation, Pennsylvania; latex peel of external mold; x1 and x2, respectively). 5, 6, Thorax of Laudonia bispinata, showing progressive change in pleural morphology down its length and herein interpreted as representing a variant of the homonomous condition (USNM 443764, from middle member of Mural Formation at Locality 61k, Mumm Peak, Mount Robson area, Alberta; x2 and x5, respectively). See text for details.

opennotspecifiedDec 2014View details →
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FIGURE 23 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 23. Bivariate scatterplots depicting intraspecific variation in various aspects of cephalic morphology of Lochmanolenellus trapezoidalis n. sp. Symbols represent phases of cephalic development. 1, Proportional ellipticity of LA. 2, Proportional length of LA. 3, Proportional length of L2. 4, Proportional length of L3. 5, Proportional length of the glabella. 6, Proportional length of the ocular lobes. 7, Proportional width of the anterior cephalic border (measured anterior to the contact of the ocular lobes with LA). 8, Proportional anterior advancement of the genal spine bases.

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FIGURE 12 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 12. Morphologically mature specimens of Lochmanolenellus pentagonalis n. sp. preserved in a compacted state in shale and siltstone. 1, Cephalon, latex peel of external mold; FMNH PE58483, x3. 2, Cephalon, latex peel of external mold; FMNH PE58484, x6. 3, Fragmentary cephalon representing largest known specimen; FMNH PE58485, x2. 4, 5, Cephalon and thorax, with enlargement of posterior portion of thorax; MCZ 110679, x4 and x8, respectively. All in dorsal view. 1–3 from collection ICS-10241, middle member of Poleta Formation, IS-S section, Montezuma Range, Esmeralda County, Nevada; 4, 5 from middle member of Mural Formation at Mumm Peak, Mount Robson area, Alberta.

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FIGURE 8 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 8. Holotype cephalon of Lochmanolenellus primus (Lochman in Cooper et al., 1952); USNM 115681. 1, Dorsal view. 2, Anterior view. 3, Left lateral view. 4, Oblique anterolateral view. All x4. From upper portion of Puerto Blanco Formation at USNM locality 801c, Cerros de la Proveedora, Caborca area, Sonora, Mexico.

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FIGURE 24 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 24. Bivariate scatterplots depicting intraspecific variation in various aspects of cephalic morphology of Lochmanolenellus trapezoidalis n. sp. Symbols represent phases of cephalic development. 1, Orientation of the distal portion of the posterior cephalic margin (distal PCM). 2, Degree of lateral flaring of the intergenal spine. 3, Proportional cephalic width, measured as the ratio of the transverse distance between the intergenal spine bases to glabella length. 4, Posterior deflection of the proximal portion of the posterior cephalic margin (proximal PCM) relative to a transverse line. 5, Degree of lateral flaring of the genal spine, quantified as the angle between the inner margin of the proximal portion of the genal spine and the distal portion of the posterior cephalic margin (distal PCM). 6, Cephalic shape, measured as the ratio of the transverse distance between the intergenal spine bases to the transverse distance between the genal spine bases.

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FIGURE 20. 1–9 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 20. 1–9, Morphologically mature cephala of Lochmanolenellus trapezoidalis n. sp. 1–4, Holotype cephalon in dorsal, anterior, right lateral, and oblique anterolateral views; GSC 137502, x3. 5, Details of prosopon on holotype cephalon; GSC 137502, x6. 6, GSC 137503, x3. 7, GSC 137504, x5. 8, 9, Fragmentary cephalon representing largest known specimen, and details of prosopon; GSC 137505, x1.5 and x3, respectively. 10–13, Hypostomes of Lochmanolenellus trapezoidalis n. sp. 10, 11, Ventral and lateral views of GSC 137506, x2. 12, GSC 137507, ventral view, x2. 13, GSC 137508, ventral view, x3. All from Unit 4 of the Rosella Formation, GSC locality 95191, Cassiar Mountains, northern British Columbia.

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FIGURE 11 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 11. Morphologically mature specimens of Lochmanolenellus pentagonalis n. sp. preserved in limestone. 1, 2, Holotype cephalon; LACMIP 12550a/b (part and latex peel of counterpart, respectively), x4. 3, Cephalon; LACMIP 12549.1, x6. 4, Cephalon, latex peel of counterpart; LACMIP 12548.5, x5. 5, Cephalon; GSC 137492, x7. 6, Cephalon with deformed right genal region, latex peel of counterpart; LACMIP 12548.1, x5. 7, Cephalon; GSC 137493, x10. 8, Cephalon; LACMIP 12551.2, x7. 9, Hypostome, ventral view; LACMIP 12551.1, x6. All in dorsal view unless specified. 1–4, 6, 8, 9 from middle member of Poleta Formation at LACMIP locality 17052, Slate Ridge, Esmeralda County, Nevada; 5 and 7 from Unit 4 of Rosella Formation, GSC locality 95190, Cassiar Mountains, northern British Columbia.

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FIGURE 19. 1 in Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia

FIGURE 19. 1, Cephalon of Lochmanolenellus cf. subquadratus 1 from middle member of Poleta Formation at LACMIP locality 26828, Magruder Mountain, Esmeralda County, Nevada; LACMIP 7401, x3. 2, Cephalon of Lochmanolenellus cf. subquadratus 2 from middle member of Poleta Formation, LACMIP locality 17039, IS-4 section, Montezuma Range, Esmeralda County, Nevada; LACMIP 12544, x7. 3, Cephalon of genus and species indet. from upper portion of Puerto Blanco Formation at USNM locality 801c, Cerros de la Proveedora, Caborca area, Sonora, Mexico; latex peel of USNM 115682, x1.5. All in dorsal view.

opennotspecifiedDec 2014View details →
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Data from: Mollusks from the upper Shackleton Limestone (Cambrian Series 2), Central Transantarctic Mountains, East Antarctica

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publicSep 2018View details →

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