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Figure 14 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 14 Radula of Theodoxus gurur Sands & Glöer sp. nov. paratypes (UGSB 20689). A. Portion of the radula showing full sets of teeth; B. Magnified view of the central and lateral teeth; C. Magnified view of the lateral and marginal teeth; D. Magnified view of the first row of marginal teeth; E. Merging point between the first and second rows of marginal teeth; F. Magnified view of the faces of inner marginal teeth belonging to the first row. Scale bars: 100 μm (A–D), 20 μm (E–F).
Figure 21 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 21 Theodoxus syriacus (Bourguignat, 1852). A–D. Specimen collected at Sultanköyü, Mardin, Turkey (UGSB 24182); E–H. Specimen from the Yanarsu Stream, Alıçlı, Turkey (UGSB 20518). I–L. Specimen collected near Başdeğirmen, Diyarbakır, Turkey (UGSB 24183); M–O; P–R. Two specimens collected by Bourguignat from İskenderun, Turkey (MHNG-MOLL-111712). All UGSB specimens incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm.
Figure 13 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 13 Theodoxus gurur Sands & Glöer sp. nov. A–D. Holotype from the Suyunbaşı spring complex, Ayrancılar, İzmir, Turkey stored in NMNL (RMNH.MOL.342197); E–G. Paratype from same locality and storage facility (RMNH.MOL.342199). Scale bars: 1 mm.
Figure 17 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 17 Theodoxus jordani (G.B. Sowerby I, 1836). A–C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakır, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus-morphotype) collected in Şanlıurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F–H; K–M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB/Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm.
Figure 10 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 10 Theodoxus fluviatilis (Linnaeus, 1758). A–D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E–H. Specimen (T. subthermalis-morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I–K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL-111736); L. Specimen (T. saulcyi-morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm.
Figure 26 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 26 Type locality of Theodoxus wilkei Sands & Glöer sp. nov. A, B. Çifteler spring, Çifteler, Eskişehir, Turkey, 39.34931°N, 31.05527°E.
Figure 20 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 20 Theodoxus pallidus (Dunker, 1861). A–D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E–H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M–P. Specimen (T. doriae-morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W–Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A–D and M–P were used in the phylogeny. Scale bars: 1 mm.
Figure 12 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 12 Type locality of Theodoxus gurur Sands & Glöer sp. nov. A, B. Suyunbaşı spring complex, Ayrancılar, İzmir, Turkey, 38.24826°N, 27.28117°E.
Figure 16 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 16 Theodoxus jordani (G.B. Sowerby I, 1836). A, B. Syntype of N. euphratica from the Euphrates River at Samawah, Iraq (ZMZ 528916); C–E. Syntype of N. michonii from Syria (MHNG-MOLL-111706); F–H. Syntype of N. karasuna collected from the Karasu River, south-eastern Turkey (ZMZ 528937); I, J; K, L. Two syntypes of N. ponsoti from Lake Muzayrīb, Syria (MNHN-IM-2000-32754); M. Syntype (labelled as paratype) of N. ponsoti from Lake Muzayrīb (MT820, RBINS); N. Syntype of N. jordani var. turris collected in the Sea of Galilee (ZMZ 528930); O, P. Syntype of N. bellardii from the Litani River, Beqaa valley, Lebanon (ZMZ 528918). Scale bars: 1 mm.
Figure 1 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 1 Terminology of key morphological characters for Theodoxus. A. Shell: a = apex, ap = aperture (im = inner margin, om = outer margin), cp = columellar plate, p = periostracum, s = spire, sh = shell height, sm = shell margin, sw = shell width, w = whorls; B. Operculum: ap = apophysis, ca = callus, cb = calcareous base, cl = conchioline lamella, la = left adductor, pa = pseudo-apophysis, ra = right adductor, rp = rib-pouch, rs = rib-shield; C. Radula: ac = A-central (c = cusp, r = ridge, tp = tooth plate), bc = B-central, cc = C-central, el = E-lateral (le = lower edge, ue = upper edge), mt1 = first row of marginal teeth (sf = small face of marginal teeth), mt2 = second row of marginal teeth, rc = R-central (ae = anterior edge, f = tooth face).
Figure 2 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 2 Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16S and ATPα (Suppl. material 1: Table S1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the "Systematic checklist" for more information). Small green (PP ≥ 0.95) and red (PP < 0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1–32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods).
Figure 28 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 28 Radula of Theodoxus wilkei Sands & Glöer sp. nov. paratypes (UGSB 20687). A. Portion of the radula showing full sets of teeth; B. Magnified view of the central and lateral teeth; C. Magnified view of the lateral and first and second rows of marginal teeth; D. Magnified view of the lateral teeth; E. The first row of marginal teeth; F. Magnified view of the faces of inner marginal teeth belonging to the first row. Scale bars: 100 μm (A–D), 20 μm (E–F).
Figure 24 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 24 Theodoxus wesselinghi Sands & Glöer sp. nov. A–D. Holotype collected in the Sakarya River, Çayköy, Bilecik, Turkey (RMNH.MOL.342200); E–G. Paratype from the same location as the holotype (RMNH.MOL.342202); H–J. Paratype from Fele, Isparta province, Turkey (RMNH.MOL.342204); K–M. Paratype from Eflatun Pınarı, Konya province, Turkey (RMNH.MOL.342206). All photographed material (A–M) is stored in NMNL. Scale bars: 1 mm.
Figure 27 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 27 Theodoxus wilkei Sands & Glöer sp. nov. A–D. Holotype (RMNH.MOL.342208); E–G. Paratype (RMNH.MOL.342210); H–J. Paratype (RMNH.MOL.342211). All photographed material is from the Çifteler Spring, Çifteler, Eskişehir, Turkey and stored in NMNL. Scale bars: 1 mm.
Figure 23 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 23 Type locality of Theodoxus wesselinghi Sands & Glöer sp. nov. A, B. Sakarya River, Çayköy, Bilecik, Turkey, 40.0439°N, 30.452°E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358°N, 31.47217°E; E, F. Eflatun Pınarı, near Sadıkhacı, Konya, Turkey, 37.8256°N, 31.6748°E; G, H. Balıkdamı Wetland spring, Eskişehir, Turkey, 39.15277°N, 31.61562°E.
Figure 25 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 25 Radula of Theodoxus wesselinghi Sands & Glöer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A–E), 20 μm (F).
Figure 3 from: Sands AF, Glöer P, Gürlek ME, Albrecht C, Neubauer TA (2020) A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96(1): 25-66. https://doi.org/10.3897/zse.96.48312
Figure 3 Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A–D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2).
FIGURE 2. Shell measurement protocol. H height L length W width. a in New taxonomic and phylogeographic data on three nominal species of the genus Septaria Férussac, 1807 (Gastropoda: Cycloneritida: Neritidae)
FIGURE 2. Shell measurement protocol. H height L length W width. a apex, ms muscle scar, s septum.
Supplementary material 2 from: Barroso CX, Pereira de Freitas JE, Matthews-Cascon H, Arruda Bezerra LE, da Cruz Lotufo TM (2020) Molecular evidences confirm the taxonomic separation of two sympatric congeneric species (Mollusca, Gastropoda, Neritidae, Neritina). ZooKeys 904: 117-130. https://doi.org/10.3897/zookeys.904.46790
Radulae of the Neritina meleagris (A) and Neritina virginea (B) analysed
Supplementary material 1 from: Barroso CX, Pereira de Freitas JE, Matthews-Cascon H, Arruda Bezerra LE, da Cruz Lotufo TM (2020) Molecular evidences confirm the taxonomic separation of two sympatric congeneric species (Mollusca, Gastropoda, Neritidae, Neritina). ZooKeys 904: 117-130. https://doi.org/10.3897/zookeys.904.46790
Alignments used to construct the phylogenetic trees and statistical parsimony network analysis
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