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Fig. 29 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 29. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1. Dermocoma sp. nov. innom. 4 from the early Albian (Early Cretaceous) of Leighton Buzzard, Great Britain; NHM EE 14844, proximal LAP. 2-5. Dermacantha pattyana gen. et sp. nov. from the Hettangian (Early Jurassic) of Fontenoille, Belgium. 2. MnhnL HE414 (holotype), proximal LAP. 3. MnhnL HE415 (paratype), proximal LAP. 4. MnhnL HE416 (paratype), median LAP. 5. MnhnL HE417 (paratype), distal LAP. 6-8. Dermacantha seeweni (Kutscher & Hary, 1991) comb. nov. from the late Sinemurian (Early Jurassic) of Bishop's Cleeve, Great Britain. 6. GZG.INV.78706, proximal LAP. 7. GZG.INV.78707, median LAP. 8. GZG.INV.78708, distal LAP. 9-11. Dermacantha leonorae gen. et sp. nov. from the early Pliensbachian (Early Jurassic) of Blockley, Great Britain. 9. GZG.INV.78710 (holotype), proximal LAP. 10. GZG.INV.78711 (paratype), median LAP. 11. GZG.INV.78712 (paratype), distal LAP. One common scale bar per species.
Fig. 36 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 36. Lateral arm plates (LAPs) of fossil and Recent ophiacanthid brittle stars in external (a) and internal (b) views. 1-3. Reitneracantha dissidens gen. et sp. nov. from the late Sinemurian to early Pliensbachian (Early Jurassic) of the Glasenbach Gorge, Austria. 1. NHMW 2012/0137/0029 (holotype), proximal LAP. 2. NHMW 2012/0137/0030 (paratype), median LAP. 3. NHMW 2012/0137/0031 (paratype), distal LAP. 4-5. Ophiohamus nanus O'Hara & Stöhr, 2006, Recent. 4. Proximal to median LAP. 5. Proximal to median LAP. 6-8. Sabinacantha archetypa gen. et sp. nov. from the late Oxfordian (Late Jurassic) of the Plettenberg, Germany. 6. GZG.INV.78771 (holotype), proximal LAP. 7. GZG.INV.78772 (paratype), median LAP. 8. GZG.INV.78773 (paratype), distal LAP. One common scale bar per species.
Fig. 8 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 8. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1. Lapidaster sp. nov. innom. 1 from the late Toarcian (Early Jurassic) of Seewen, Switzerland; NHMB M11208, proximal to median LAP. 2-4. Lapidaster wolfii gen. et sp. nov. from the Bajocian-Bathonian boundary (Middle Jurassic) of Touert, France. 2. GZG.INV.78508 (holotype), proximal LAP. 3. GZG. INV.78509 (paratype), median LAP. 4. GZG.INV.78510 (paratype), distal LAP. 5-7. Lapidaster varuna gen. et sp. nov. from the Callovian of Jumara, India. 5. GZG.INV.78511 (holotype), proximal LAP. 6. GZG.INV.78512 (paratype), median LAP. 7. GZG.INV.78513 (paratype), distal LAP. One common scale bar per species is given.
Fig. 24 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 24. Fossil lateral arm plates in external (a) and internal (b) views and articulated arm fragment of ophiacanthid brittle stars. 1-2. Alternacantha arges sp. nov. from the late Oxfordian (Late Jurassic) of Raedersdorf, France; M 610 in dorsal (1a) and ventral (1b) views and with details of dorsal (2a) and ventral (2b) arm plating. 3-6. Alternacantha dilionessa sp. nov. from the early Kimmeridgian of the Pointe du Chay, France. 3. GZG.INV.78662 (holotype), proximal LAP. 4. GZG.INV.78663 (paratype), proximal LAP. 5. GZG.INV.78664 (paratype), median LAP. 6. GZG.INV.78665 (paratype), distal LAP. One common scale bar for 1, for 2 and for 3-6, respectively.
Fig. 7 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 7. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-3. Lapidaster caeloscopus gen. et sp. nov. from the late Sinemurian to early Pliensbachian (Early Jurassic) of the Glasenbach Gorge, Austria. 1. NHMW 2012/0137/0005 (holotype), proximal LAP. 2. NHMW 2012/0137/0006 (paratype), median LAP. 3. NHMW 2012/0137/0007 (paratype), distal LAP. 4-6. Lapidaster hystricarboris gen. et sp. nov. from the early Pliensbachian (Early Jurassic) of Blockley, Great Britain. 4. GZG.INV.78501 (holotype), proximal LAP. 5. GZG.INV.78502 (paratype), median LAP. 6. GZG.INV.78503 (paratype), distal LAP. 7-8. Lapidaster fasciatus (Kutscher & Villier, 2003) comb. nov. from the middle Toarcian (Early Jurassic) of Le Clapier, France. 7. GZG.INV.78505, proximal LAP. 8. GZG.INV.78506, distal LAP. One common scale bar per species is given.
Fig. 18 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 18. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-2. Ophiotreta sp. nov. innom 2 from the Langhian (middle Miocene) of Stotzing (1) and Mannersdorf (2), Austria. 1. NHMW 2012/0139/0003, proximal LAP. 2. NHMW 2012/0139/0001, proximal LAP. 3-4. Europacantha paciphila gen. et sp. nov. from the late Anisian (Middle Triassic) of Felsöörs, Hungary. 3. MHI 2090/1 (holotype), proximal to median LAP. 4. MHI 2091/1 (paratype), proximal LAP. 5-7. Ophiacantha jaegeri sp. nov. from the latest Hauterivian (Early Cretaceous) of Sarstedt, Germany. 5. GZG.INV.78595 (holotype), proximal LAP. 6. GZG.INV.78596 (paratype), median LAP. 7. GZG. INV.78597 (paratype), distal LAP. 8-9. Ophiacantha sp. nov. innom 1 from the latest Aptian to earliest Albian (Early Cretaceous) of Blake Nose, NE Atlantic. 8. GZG.INV.78599, proximal LAP. 9. GZG. INV.78600, distal LAP. 10. Ophiacantha sp. nov. innom 2 from the latest Aptian to earliest Albian (Early Cretaceous) of Blake Nose, NE Atlantic; GZG.INV.78601, distal LAP. One common scale bar per species.
Fig. 5 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 5. Diversity of ophiacanthid brittle stars, given as species richness per 5 Ma intervals, in shallowwater (<200 m palaeodepth) environments through time, together with corresponding numbers of samples and of lateral arm plates (LAPs) counted (both ophiacanthids and non-ophiacanthids) per interval. Note that the scale for specimen numbers is increased by factor 103.
Fig. 3. A in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 3. A. Evolutionary tree of the Ophiacanthidae based on the reweighed dataset tree of Fig. 2B. Calibration is based on nodes marked as 1 (oldest occurrence of small tentacle pores, Europacantha paciphila sp. nov. from the Anisian of Hungary) and 2 (oldest occurrence of lateral arm plate morphology exclusively found in the Ophiomitra-Ophiocamax clade, Reitneracantha dissidens sp. nov. from the late Sinemurian to early Pliensbachian of Austria), and on the first occurrences of a number of extinct genera (Leadagmara Thuy et al., 2012, Dermocoma Hess, 1964, Inexpectacantha Thuy, 2011). Thick lines indicate the observed fossil record of the actual genus, doted lines denote the fossil record of a genus within the same lineage (e.g. Lapidaster sp. nov. in Ophiologimus H.L. Clark, 1911), and thin lines are the expected ranges. B. Relative abundances of the Ophiacanthidae in shallow (<200 m palaeodepth, small dots) and deep (> 200 m palaeodepth, asterisks) environments through time.
Fig. 33 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 33. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-3. Inexpectacantha ritae sp. nov. from the Hettangian (Early Jurassic) of Bourglinster, Luxembourg. 1. MnhnL HE419 (holotype), proximal LAP. 2. MnhnL HE420 (paratype), median LAP. 3. MnhnL HE421 (paratype), distal LAP. 4-6. Inexpectacantha weisi sp. nov. from the Hettangian (Early Jurassic) of Fontenoille, Belgium. 4. MnhnL HE425 (holotype), proximal LAP. 5. MnhnL HE426 (paratype), median LAP. 6. MnhnL HE427 (paratype), distal LAP. 7-8. Inexpectacantha lunaris (Hess, 1962) comb. nov. from the late Sinemurian (Early Jurassic) of Bishop's Cleeve, Great Britain. 7. GZG. INV.78752, proximal LAP. 8. GZG.INV.78753, proximal to median LAP. 9-10. Inexpectacantha acrobatica Thuy, 2011 from the early Pliensbachian (Early Jurassic) of Blockley, Great Britain (9) and the early Toarcian (Early Jurassic) of Aix-sur-Cloie, Belgium (10). 9. GZG.INV.78755, proximal LAP. 10. GZG.INV.78757, juvenile proximal LAP. One common scale bar per species.
Fig. 32 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 32. Fossil skeletal plates of ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views. 1. Ophiomitrella? sp. 2 from the Bathonian (Middle Jurassic) of Jumara, India; GZG. INV.78740, median to distal LAP. 2. Ophiomitrella sp. nov. from the late Maastrichtian (Late Cretaceous) of Maastricht, the Netherlands; NHMM JJ 5104, proximal LAP. 3-6. Ophiomalleus beneficarum gen. et sp. nov. from the Callovian of Bauer-Wehrland (3), Althüttendorf (4-5) and Hohensaaten (6), Germany. 3. GZG.INV.78741 (holotype), proximal to median LAP. 4. GZG.INV.78742 (paratype), proximal LAP. 5. GZG.INV.78743 (paratype), distal LAP. 6. GZG.INV.78744 (paratype), arm spine. 7-9. Ophiomalleus stevenwilsoni gen. et sp. nov. from the latest Hauterivian (Early Cretaceous) of Sarstedt, Germany. 7. GZG.INV.78748 (holotype), proximal LAP. 8. GZG.INV.78749 (paratype), median LAP. 9. GZG. INV.78750 (paratype), distal LAP. One common scale bar per species except for 6.
Fig. 28 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 28. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-4. Dermocoma numbergerorum sp. nov. from the middle Oxfordian (Late Jurassic) of Foug, France. 1. GZG.INV.78695 (holotype), proximal LAP. 2. GZG.INV.78696 (paratype), proximal LAP. 3. GZG.INV.78697 (paratype), median LAP. 4. GZG.INV.78698 (paratype), distal LAP. 5-8. Dermocoma simonschneideri sp. nov. from the early Kimmeridgian (Late Jurassic) of the Pointe du Chay, France (5) and the late Kimmeridgian (Late Jurassic) of Trancoso, Portugal (6-8). 5. GZG.INV.78700 (holotype), proximal LAP. 6. GZG.INV.78701 (paratype), proximal LAP. 7. GZG.INV.78702 (paratype), median LAP. 8. GZG. INV.78703 (paratype), distal LAP. 9. Dermocoma sp. nov. innom. 2 from the early Kimmeridgian (Late Jurassic) of Geisingen, Germany; GZG.INV.78705, median to distal LAP. 10-11. Dermocoma sp. nov. innom. 3 from the late Valanginian (Early Cretaceous) of the Ternberg Nappe, Austria. 10. NHMW 2012/0138/0009, proximal LAP. 11. NHMW 2012/0138/0010, distal LAP. One common scale bar per species.
Fig. 2 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 2. Phylogeny of the Ophiacanthidae. A. Majority rule consensus of 2572 most parsimonious trees (182 steps length) resulting from the phylogenetic analysis of the full data set. B. Strict consensus of nine most parsimonious trees (47.91375 steps length) resulting from the analysis of the reweighed dataset (characters reweighed by their maximum rescaled consistency index). Numbers above branches represent bootstrap support for the respective nodes (10 000 replicates); numbers below branches are node frequencies of the majority rule consensus tree.
Fig. 27 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 27. Fossil lateral arm plates in external (a) and internal (b) views and articulated arm fragment of ophiacanthid brittle stars. 1-2. Dermocoma wrighti Hess, 1964 from the early Callovian (Middle Jurassic) of Liesberg, Switzerland. 1. NHMB M11218, proximal LAP. 2. NHMB M11219, distal LAP. 3-4. Dermocoma sp. 1 from the Callovian (Middle Jurassic) of Jumara, India. 3. GZG.INV.78687, proximal LAP. 4. GZG.INV.78688, distal LAP. 5-8. Dermocoma biformis (Hess, 1975) comb. nov. from the late Oxfordian (Late Jurassic) of Guldental, Switzerland (5-6) and Savigna, France (7-8). 5. NHMB M11220, proximal LAP. 6. NHMB M11221, distal LAP. 7. GZG.INV.78690, proximal LAP. 8. GZG. INV.78692, proximal arm fragment in ventral (a) and dorsal (b) views. One common scale bar for 1-2, 3-4, 5-6, 7 and 8 respectively.
Fig. 1 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 1. Ophiacanthid brittle star lineages and their fossil record in shallow (<200 m palaeodepth, thick grey lines) and deep (> 200 m palaeodepth, thick black lines) environments through time. Extant lineages are indicated by the thin median line extending to the Holocene.
Fig. 6 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 6. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1. Eolaxoporus sp. from the latest Anisian (Middle Triassic) of Oberscheffach, Germany, MHI 2083/1, median LAP. 2-4. Eolaxoporus hagdorni gen. et sp. nov. from the late Carnian (Late Triassic) of Jushui, China. 2. MHI 2084/1 (holotype), proximal LAP. 3. MHI 2085/1 (paratype), median LAP. 4. MHI 2086/1 (paratype), distal LAP. 5. Eolaxoporus sp. nov. innom. from the early Carnian (Late Triassic) of Milieres, Italy; GZG.INV.78500, proximal to median LAP. 6-8. Eolaxoporus imminens gen. et sp. nov. from the late Sinemurian to early Pliensbachian (Early Jurassic) of the Glasenbach Gorge, Austria. 6. NHMW 2012/0137/0001 (holotype), proximal LAP. 7. NHMW 2012/0137/0002 (paratype), median LAP. 8. NHMW 2012/0137/0003 (paratype), distal LAP. One common scale bar per species is given.
Fig. 23 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 23. Fossil lateral arm plates in external (a) and internal (b) views and articulated arm fragments of Alternacantha schwermannorum sp. nov. from the late Oxfordian (Late Jurassic) of Savigna, France. 1. GZG.INV.78654 (holotype), proximal LAP. 2. GZG.INV.78655 (paratype), median LAP. 3. GZG. INV.78656 (paratype), distal LAP. 4. GZG.INV.78657 (paratype), proximal arm fragment in ventral (a) and dorsal (b) views and with detail in dorso-lateral (c) view. 5. GZG.INV.78658 (paratype), distal arm fragment in dorsal (a) and ventral (b) view. One common scale bar for 1-3.
Appendix 1 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Appendix 1. Faunal list for the assemblages studied with corresponding counts of lateral arm plates or articulated individual (art.). Note that previously published assemblages which lack ophiacanthids or for which no significant taxonomic changes or new counts are proposed have been omitted. Records marked with an asterisk (*) are not described in the present study.
Fig. 35 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 35. Skeletal plates and arm fragments of fossil and Recent ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views. 1-3. Ophiocamax vitrea Lyman, 1878, Recent. 1. Proximal LAP. 2. Median LAP. 3. Dorsal arm plate. 4-5. Ophiocamax hystrix Lyman, 1878, Recent. 4. Proximal LAP. 5. Median LAP. 6. Ophiocamax austera Verrill, 1899, Recent; proximal LAP. 7-11. Ophiocamax dorotheae sp. nov. from the late Oxfordian (Late Jurassic) of Savigna, France. 7. GZG.INV.78763 (holotype), proximal LAP. 8. GZG.INV.78764 (paratype), median LAP. 9. GZG.INV.78765 (paratype), distal LAP. 10. GZG.INV.78766 (paratype), proximal arm fragment in ventral view. 11. GZG.INV.78767 (paratype), median arm fragment in dorsal view. One common scale bar per species except for 10 and 11.
Fig. 17 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 17. Fossil skeletal plates of ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views. 1-2. Ophiotreta striata (Kutscher & Jagt, 2000) comb. nov. from the early Maastrichtian of Rügen, Germany. 1. GZG.INV.78588, proximal LAP. 2. GZG.INV.78589, distal LAP. 3-11. Ophiotreta dendrophyllicola sp. nov. from the middle Danian (Paleocene) of Fakse, Denmark. 3. MGUH 30236 (holotype), proximal LAP. 4. NHMM 2012 050 (paratype), median LAP. 5. NHMM 2012 051 (paratype), distal LAP. 6. NHMM 2012 052 (paratype), vertebra in distal view. 7. NHMM 2012 053 (paratype), vertebra in proximal view. 8. NHMM 2012 054 (paratype), vertebra in dorsal view. 9. NHMM 2012 055 (paratype), oral plate in adradial (a) and abradial (b) view. 10. NHMM 2012 056 (paratype), arm spine fragment. 11. NHMM 2012 057 (paratype), arm spine fragment. 12-15. Ophiotreta hedone sp. nov. from the middle Lutetian (Eocene) of Grignon, France. 12. GZG.INV.78590 (holotype), proximal LAP. 13. GZG.INV.78591 (paratype), median LAP. 14. GZG.INV.78592 (paratype), distal LAP. 15. GZG.INV.78593 (paratype), median LAP. One common scale bar per species.
Fig. 4 in Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)
Fig. 4. Palaeogeographic reconstruction for the Middle Triassic [after Smith et al. (1994)] with positions of currently known Triassic ophiuroid occurrences (grey areas indicate emerged land). Round dots indicate assemblages quantitatively assessed in this study (see Table 1 for details). 1. Fischerwiese, Oberscheffach, Schillingstadt. 2. Górazdze, Strzelce Opolski, Felsöörs, Sóly. 3. Alpe di Specie, Romerlo, Milieres, Recoaro. 4. Kitakami Mountains. 5. Jushui. Squares indicate previously published Triassic records. At the same positions as round dots 1-3: Bachmayer & Kollmann (1968), Broglio Loriga & Berti Cavicchi (1972), Hess (1970a), Kutscher (1987b, 2000), Radwański (2002), Salamon (2004) and ophiuroid records revised by Hess (1965b). Squares: 6. Calzada & Gutiérrez (1988). 7. Hess (1972b), Kristan-Tollmann et al. (1979). 8. Twitchett et al. (2005). 9. Zonneveld (2001). 10. Kummel & Teichert (1970). 11. Runnegar (1969). 12. Kristan-Tollmann & Gramann (1992). At the same position as round dot 5: Yang (1960), Feng (1985), Chen et al. (2004). Square 13. Ishida et al. (2011).
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