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FIG. 30. Heteroctenus Pocock, 1893 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 30. Heteroctenus Pocock, 1893, metasoma and telson, lateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. H. junceus (Herbst, 1800), ♂ (AMNH). E. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). Scale bars = 5 mm.

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FIG. 27. Heteroctenus Pocock, 1893 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 27. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. junceus (Herbst, 1800), comb. nov., ♂ (AMNH). B. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. H. junceus (Herbst, 1800), ♀ (AMNH). D. H. princeps (Karsch, 1879), comb. nov., ♀ (AMNH). Scale bars = 2 mm.

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FIG. 24 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A–D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E–H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I–M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N–R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S–U. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm.

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FIG. 14 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm.

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FIG. 20 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm.

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FIG. 23 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 23. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (D, L, R), ectal aspect (A, F, H, O, T), ectal aspect, detail (B, K, M, V), ental aspect (G, I, P, S) and ental aspect, detail (C, J, N, Q, U). A–D. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). E–G. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). H–K. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). L–N. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). O–R. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). S–V. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). Scale bars = 1 mm.

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FIG. 9 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses).

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FIG. 17 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm.

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FIG. 6 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 6. Map of northern South America, plotting known locality records of Jaguajir pintoi (Mello-Leitão, 1932), comb. nov. (circles), Rhopalurus caribensis Teruel and Roncallo, 2008 (pentagons), and Rhopalurus ochoai, sp. nov. (triangles).

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FIG. 1 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil.

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FIG. 3 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 3. Map of Cuba and neighboring islets, plotting known locality records of three species of Heteroctenus Pocock, 1893: A. H. junceus (Herbst, 1800) (circles). B. H. garridoi (Armas, 1974), comb. nov. (squares); H. gibarae (Teruel, 2006), comb. nov. (crosses).

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FIG. 26. Heteroctenus Pocock, 1893 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 26. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♀ (AMNH). B. H. bonettii (Armas and Marcano Fondeur, 1987), comb. nov., ♀ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♀ (AMNH). Scale bars = 2 mm.

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FIG. 12 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP).

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FIG. 5 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 5. Map of northern South America, plotting known locality records of Rhopalurus laticauda Thorell, 1876 (squares).

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FIG. 22 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm.

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FIG. 19 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 19. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♀ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm.

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FIG. 2 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004.

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FIG. 11 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP).

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FIG. 25. Heteroctenus Pocock, 1893 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 25. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). Scale bars = 2 mm.

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FIG. 18 in Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)

FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm.

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