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Text-fig. 6. Scanning electron micrographs of multicarpellate and apocarpous floral structures from the Early Cretaceous Puddledock locality, Virginia, USA (a, b: PP43701, Puddledock sample 001; c: PP43000x, Puddledock sample 073). a) Anacostia? sp., strongly compressed, elongated receptacle with spirally arranged carpels (red dots; not all shown); note larger size compared to the other floral structures; b) Numerous Anacostia type pollen grains in proximal view from the base of floral structure in (a); note graded reticulum over the proximal pole of the pollen grains; c) Elongated receptacle with numerous carpels in a spiral arrangement, possibly representing an earlier developmental stage of Anacostia? sp. Scale bars = 1 mm (a, c), 10 µm (b). in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal

Text-fig. 6. Scanning electron micrographs of multicarpellate and apocarpous floral structures from the Early Cretaceous Puddledock locality, Virginia, USA (a, b: PP43701, Puddledock sample 001; c: PP43000x, Puddledock sample 073). a) Anacostia? sp., strongly compressed, elongated receptacle with spirally arranged carpels (red dots; not all shown); note larger size compared to the other floral structures; b) Numerous Anacostia type pollen grains in proximal view from the base of floral structure in (a); note graded reticulum over the proximal pole of the pollen grains; c) Elongated receptacle with numerous carpels in a spiral arrangement, possibly representing an earlier developmental stage of Anacostia? sp. Scale bars = 1 mm (a, c), 10 µm (b).

Text-fig. 9. Number of required character state changes under parsimony (steps) for various positions of Atlantocarpus virginiensis, based on the Doyle and Endress character matrix and backbone tree (Doyle and Endress 2000, 2014). in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal

Text-fig. 9. Number of required character state changes under parsimony (steps) for various positions of Atlantocarpus virginiensis, based on the Doyle and Endress character matrix and backbone tree (Doyle and Endress 2000, 2014).

Text-fig. 5. Scanning electron micrographs (d–g) and synchrotron radiation X-ray tomographic microscopy orthoslices (a–c) of flowers of Atlantocarpus virginiensis gen. et sp. nov. (a–d: holotype, PP43780, Puddledock sample 156), Atlantocarpus sp. from the Early Cretaceous Buarcos locality (e, f: S105025, Buarcos sample 244) and receptacle of Atlantocarpus? from the Early Cretaceous Vale de Água locality (g: S101300, Vale de Água sample 141). a) Flower in lateral view showing scar from a single bract (br), attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels; b) Flower in lateral view showing expanded basal portion of the elongated receptacle and young carpels; c) Flower in longitudinal section showing expanded basal portion of the elongated receptacle and young carpels; note the irregular, possibly expanded stigmatic region (arrow heads), (orthoslice yz0340); d) Flower in lateral view showing attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels with possible grooved stigmatic regions (arrow heads); e) Flower in lateral view showing expanded basal portion of elongated receptacle and young carpels; f) Detail of flower in (e) showing in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal

Text-fig. 5. Scanning electron micrographs (d–g) and synchrotron radiation X-ray tomographic microscopy orthoslices (a–c) of flowers of Atlantocarpus virginiensis gen. et sp. nov. (a–d: holotype, PP43780, Puddledock sample 156), Atlantocarpus sp. from the Early Cretaceous Buarcos locality (e, f: S105025, Buarcos sample 244) and receptacle of Atlantocarpus? from the Early Cretaceous Vale de Água locality (g: S101300, Vale de Água sample 141). a) Flower in lateral view showing scar from a single bract (br), attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels; b) Flower in lateral view showing expanded basal portion of the elongated receptacle and young carpels; c) Flower in longitudinal section showing expanded basal portion of the elongated receptacle and young carpels; note the irregular, possibly expanded stigmatic region (arrow heads), (orthoslice yz0340); d) Flower in lateral view showing attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels with possible grooved stigmatic regions (arrow heads); e) Flower in lateral view showing expanded basal portion of elongated receptacle and young carpels; f) Detail of flower in (e) showing

Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy orthoslices of flowers of Lambertiflora virginiense gen. et sp. nov. from the Early Cretaceous Puddledock locality, Virginia, USA (holotype, PP53796, Puddledock sample 081). White dots – tepals, yellow dots – stamens or staminodes, red dot – central conical gynoecial region. a) Flower in longitudinal section showing elongated overlapping tepals, remains of probable poorly developed stamens or staminodes and probable poorly developed carpels on the central conical gynoecial region of the receptacle (orthoslice yz0454); b) Flower in transverse section showing rhomboidal bases of 30 tepals, nine poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1160); c) Flower in transverse section at the level of the floral receptacle showing 30 tepals, nine of the poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1250). Scale bars = 250 µm (a–c). in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal

Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy orthoslices of flowers of Lambertiflora virginiense gen. et sp. nov. from the Early Cretaceous Puddledock locality, Virginia, USA (holotype, PP53796, Puddledock sample 081). White dots – tepals, yellow dots – stamens or staminodes, red dot – central conical gynoecial region. a) Flower in longitudinal section showing elongated overlapping tepals, remains of probable poorly developed stamens or staminodes and probable poorly developed carpels on the central conical gynoecial region of the receptacle (orthoslice yz0454); b) Flower in transverse section showing rhomboidal bases of 30 tepals, nine poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1160); c) Flower in transverse section at the level of the floral receptacle showing 30 tepals, nine of the poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1250). Scale bars = 250 µm (a–c).

Text-fig. 54. Scanning electron microscope (SEM; a–e, g, h) and synchrotron x-ray microscopy (SRXTM; f, i) images of unnamed angiosperm fruits and seeds; Torres Vedras locality, Portugal. a) Follicle sp. 1, narrow elongate follicle with sessile and decurrent stigma; b) Follicle sp. 2, broad, dehisced follicle with elongate and transverse fibers lining the locule; c) Angiosperm seed sp. 1 with thin, smooth seed coat; d) Angiosperm seed sp. 2 with verrucate seed coat; e) Angiosperm seed sp. 3 with exotestal and foveolate seed coat; f) Angiosperm seed sp. 4 with smooth seed coat; g, h) Angiosperm seed sp. 5 with raised epidermal cells forming a reticulate pattern; i) Wedge-shaped angiosperm fruit with remains of floral parts near apex. Specimens, TV43-S136726 (a), TV44-S148144 (b), TV43-S170074 (c), TV43-S136747 (d), TV43-S170073 (e), TV38-S174615 (f), TV44-S148003 (g), TV44-S148004 (h), TV43-S174685 (i). Scale bars 300 Μm (a–i). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 54. Scanning electron microscope (SEM; a–e, g, h) and synchrotron x-ray microscopy (SRXTM; f, i) images of unnamed angiosperm fruits and seeds; Torres Vedras locality, Portugal. a) Follicle sp. 1, narrow elongate follicle with sessile and decurrent stigma; b) Follicle sp. 2, broad, dehisced follicle with elongate and transverse fibers lining the locule; c) Angiosperm seed sp. 1 with thin, smooth seed coat; d) Angiosperm seed sp. 2 with verrucate seed coat; e) Angiosperm seed sp. 3 with exotestal and foveolate seed coat; f) Angiosperm seed sp. 4 with smooth seed coat; g, h) Angiosperm seed sp. 5 with raised epidermal cells forming a reticulate pattern; i) Wedge-shaped angiosperm fruit with remains of floral parts near apex. Specimens, TV43-S136726 (a), TV44-S148144 (b), TV43-S170074 (c), TV43-S136747 (d), TV43-S170073 (e), TV38-S174615 (f), TV44-S148003 (g), TV44-S148004 (h), TV43-S174685 (i). Scale bars 300 Μm (a–i).

Text-fig. 53. Scanning electron microscope (SEM) images of tricolpate pollen of Nicholsia brevicolpites gen. et sp. nov. from a coprolite containing several kinds of pollen; Torres Vedras locality, Portugal. a) Detail of the coprolite that yielded the tricolpate pollen in this Text-figure showing several different kinds of pollen grains; b–d) Tricolpate pollen grains in polar (b) and equatorial (c, d) views showing colpi, coarsely granular aperture membrane and foveolate tectum. Specimen, TV44-S137906-03 (holotype; a–d). Scale bars 30 Μm (a), 6 Μm (b–d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 53. Scanning electron microscope (SEM) images of tricolpate pollen of Nicholsia brevicolpites gen. et sp. nov. from a coprolite containing several kinds of pollen; Torres Vedras locality, Portugal. a) Detail of the coprolite that yielded the tricolpate pollen in this Text-figure showing several different kinds of pollen grains; b–d) Tricolpate pollen grains in polar (b) and equatorial (c, d) views showing colpi, coarsely granular aperture membrane and foveolate tectum. Specimen, TV44-S137906-03 (holotype; a–d). Scale bars 30 Μm (a), 6 Μm (b–d).

Text-fig. 50. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of a flower and mature carpels of Reyanthus lusitanicus gen. et sp. nov.; Torres Vedras locality, Portugal. a) Holotype; multicarpellate flower showing numerous carpels surrounded by remains of the androecium and perianth; note the bract subtending the flower (arrowhead); b, c) Longitudinal sections perpendicular to section in (f) (b, SRXTM orthoslice xz0461) and (c, SRXTM cut voltex xz0540-0580) showing receptacle, carpels and remains of the androecium and perianth; note prominent cavities formed by the oil cells, subtending bract (arrow) and bract subtending fragmentary bud (arrowhead); d, e) Apical (d) and lateral (e) views of gynoecium showing laterally flattened carpels and dorsi-ventrally flattened stamen or staminode (* in e); f) Longitudinal section (SRXTM orthoslice yz0405) of gynoecium showing conical receptacle and carpels with densely spaced oil cells; g, h) Mature carpel of Reyanthus lusitanicus gen. et sp. nov. showing slightly curved form, rounded apex and base that tapers toward the attachment to the receptacle; i) Surface of carpel showing embedded oil cells; j) Transverse section through three carpels showing attachment of developing ovules near the ventral sutures and oil cells in the wall just below the surface (SRXTM cut voltex xy0770-0845). Specimens TV299-S136716 (holotype; a–f), TV299-S136717 (g–j). Scale bars 300 Μm (a–d, f–h), 150 Μm (e, j), 30 Μm (i). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 50. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of a flower and mature carpels of Reyanthus lusitanicus gen. et sp. nov.; Torres Vedras locality, Portugal. a) Holotype; multicarpellate flower showing numerous carpels surrounded by remains of the androecium and perianth; note the bract subtending the flower (arrowhead); b, c) Longitudinal sections perpendicular to section in (f) (b, SRXTM orthoslice xz0461) and (c, SRXTM cut voltex xz0540-0580) showing receptacle, carpels and remains of the androecium and perianth; note prominent cavities formed by the oil cells, subtending bract (arrow) and bract subtending fragmentary bud (arrowhead); d, e) Apical (d) and lateral (e) views of gynoecium showing laterally flattened carpels and dorsi-ventrally flattened stamen or staminode (* in e); f) Longitudinal section (SRXTM orthoslice yz0405) of gynoecium showing conical receptacle and carpels with densely spaced oil cells; g, h) Mature carpel of Reyanthus lusitanicus gen. et sp. nov. showing slightly curved form, rounded apex and base that tapers toward the attachment to the receptacle; i) Surface of carpel showing embedded oil cells; j) Transverse section through three carpels showing attachment of developing ovules near the ventral sutures and oil cells in the wall just below the surface (SRXTM cut voltex xy0770-0845). Specimens TV299-S136716 (holotype; a–f), TV299-S136717 (g–j). Scale bars 300 Μm (a–d, f–h), 150 Μm (e, j), 30 Μm (i).

Text-fig. 47. Scanning electron microscope (SEM) images of reticulate-foveolate pollen on the surface of a fruit of Serialis antiquum; Torres Vedras locality, Portugal. a, b) Pollen grain showing angular outline and tectum perforated by foveolae of different sizes; note the braided tectum ornamentation formed by aggregations of small ridges (b); aperture not known; c, d) Pollen grain showing angular outline, tectum perforated by foveolae that become smaller toward the presumed pole; note the relatively smooth tectum and faint pattern of braiding ridges (d); aperture not known. Specimens, TV43-S171535-01 (a, b), TV43-S171535-02 (c, d). Scale bars 6 Μm (a, c), 1 Μm (b, d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 47. Scanning electron microscope (SEM) images of reticulate-foveolate pollen on the surface of a fruit of Serialis antiquum; Torres Vedras locality, Portugal. a, b) Pollen grain showing angular outline and tectum perforated by foveolae of different sizes; note the braided tectum ornamentation formed by aggregations of small ridges (b); aperture not known; c, d) Pollen grain showing angular outline, tectum perforated by foveolae that become smaller toward the presumed pole; note the relatively smooth tectum and faint pattern of braiding ridges (d); aperture not known. Specimens, TV43-S171535-01 (a, b), TV43-S171535-02 (c, d). Scale bars 6 Μm (a, c), 1 Μm (b, d).

Text-fig. 48. Scanning electron microscope (SEM) images of monocolpate pollen from a coprolite; Torres Vedras locality, Portugal. a) Coprolite that yielded the pollen in this Text-figure as well as several other kinds of pollen; b) Reticulum of pollen grain showing the tall, sharp muri and obconical columellae; c–e) Grains of Goczania inaequalis sp. nov. intermixed with pollen grains that have a coarse, loosely attached reticulum with tall, sharp muri and sparse columellae; note the weak transverse striations on the muri and that the reticulum is much larger than the main body of the grain that it encloses. Specimen, TV44-S148023. Scale bars 300 Μm (a), 6 Μm (c–e), 3 Μm (b). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 48. Scanning electron microscope (SEM) images of monocolpate pollen from a coprolite; Torres Vedras locality, Portugal. a) Coprolite that yielded the pollen in this Text-figure as well as several other kinds of pollen; b) Reticulum of pollen grain showing the tall, sharp muri and obconical columellae; c–e) Grains of Goczania inaequalis sp. nov. intermixed with pollen grains that have a coarse, loosely attached reticulum with tall, sharp muri and sparse columellae; note the weak transverse striations on the muri and that the reticulum is much larger than the main body of the grain that it encloses. Specimen, TV44-S148023. Scale bars 300 Μm (a), 6 Μm (c–e), 3 Μm (b).

Text-fig. 45. Scanning electron microscope (SEM) images of monocolpate pollen of Dinisia portugallica gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Two pollen grains showing poorly defined distal aperture (arrowhead) and distinctive vermiform reticulum forming luminae of variable shapes and sizes; note especially the irregularly and incomplete reticulum in the grain on the left; c) Reticulum showing smooth, vermiform muri attached to the smooth surface of the foot layer by long columellae; note that columellae often terminate segments of muri that are not closed; d, e) Pollen grains showing proximal surface (d), poorly defined distal aperture (e, arrowhead) and distinctive vermiform reticulum supported by long columellae; note dense covering of small, spherical orbicules on the inner surface of the anther wall. Specimen, TV44-S148216 (holotype). Scale bars 300 Μm (a), 6 Μm (b, d, e), 3 Μm (c). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 45. Scanning electron microscope (SEM) images of monocolpate pollen of Dinisia portugallica gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Two pollen grains showing poorly defined distal aperture (arrowhead) and distinctive vermiform reticulum forming luminae of variable shapes and sizes; note especially the irregularly and incomplete reticulum in the grain on the left; c) Reticulum showing smooth, vermiform muri attached to the smooth surface of the foot layer by long columellae; note that columellae often terminate segments of muri that are not closed; d, e) Pollen grains showing proximal surface (d), poorly defined distal aperture (e, arrowhead) and distinctive vermiform reticulum supported by long columellae; note dense covering of small, spherical orbicules on the inner surface of the anther wall. Specimen, TV44-S148216 (holotype). Scale bars 300 Μm (a), 6 Μm (b, d, e), 3 Μm (c).

Text-fig. 43. Scanning electron microscope (SEM) images of monocolpate pollen of Kempia longicolpites gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Pollen grains showing the very long distal aperture and loosely attached reticulum; c, d) Pollen grains showing the very long, clearly defined distal aperture that extends around the ends of the grain; e) Reticulum in nonapertural region showing the smooth muri loosely attached to the smooth surface of the foot layer by short columellae; f) Internal view of reticulum showing the short columellae loosened from the foot layer. Specimen, TV44-S105018 (holotype). Scale bars 300 Μm (a), 30 Μm (b), 6 Μm (c, d), 3 Μm (e, f). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 43. Scanning electron microscope (SEM) images of monocolpate pollen of Kempia longicolpites gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Pollen grains showing the very long distal aperture and loosely attached reticulum; c, d) Pollen grains showing the very long, clearly defined distal aperture that extends around the ends of the grain; e) Reticulum in nonapertural region showing the smooth muri loosely attached to the smooth surface of the foot layer by short columellae; f) Internal view of reticulum showing the short columellae loosened from the foot layer. Specimen, TV44-S105018 (holotype). Scale bars 300 Μm (a), 30 Μm (b), 6 Μm (c, d), 3 Μm (e, f).

Text-fig. 46. Scanning electron microscope (SEM) images of monocolpate pollen of Teebacia hughesii gen. et sp. nov. pollen from a stamen fragment; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure (d, e); b) Detail of detached reticulum showing inner surface of muri and scattered columellae; note the finely granular covering of the muri and columellae; c, d) Detail of reticulum showing the outer surface of muri with supratectal ornamentation of narrow ridges; note orbicules attached to the reticulum (d); e, f) Pollen grains showing loose, beaded, reticulum with long columellae; note continuous muri bordering the apertures and densely spaced minute orbicules lining the inner surface of the anther wall (e, arrowheads). Specimens, TV44-S136666 (holotype; a, d, e), TV44-S149207 (b, c, f). Scale bars 300 Μm (a), 6 Μm (e, f), 3 Μm (b), 1.5 Μm (c), 1.2 Μm (d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 46. Scanning electron microscope (SEM) images of monocolpate pollen of Teebacia hughesii gen. et sp. nov. pollen from a stamen fragment; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure (d, e); b) Detail of detached reticulum showing inner surface of muri and scattered columellae; note the finely granular covering of the muri and columellae; c, d) Detail of reticulum showing the outer surface of muri with supratectal ornamentation of narrow ridges; note orbicules attached to the reticulum (d); e, f) Pollen grains showing loose, beaded, reticulum with long columellae; note continuous muri bordering the apertures and densely spaced minute orbicules lining the inner surface of the anther wall (e, arrowheads). Specimens, TV44-S136666 (holotype; a, d, e), TV44-S149207 (b, c, f). Scale bars 300 Μm (a), 6 Μm (e, f), 3 Μm (b), 1.5 Μm (c), 1.2 Μm (d).

Text-fig. 42. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis sp. 1 (a–c) from an isolated anther and Piercipollis sp. 2 (d–f) from a coprolite (not shown); Torres Vedras locality, Portugal. a) Narrow elongate anther that yielded the pollen in (b) and (c); b) Distal view of pollen grain showing well-developed reticulum that is only loosely attached to the foot layer; note the smooth foot layer (arrowhead) of a grain from which the reticulum has become detached; c) Reticulum showing the smooth muri and long columellae that are mostly detached from the foot layer; d–f) Pollen grains in distal (d, e), and lateral views (f) showing the very long colpus and the well-developed reticulum only loosely attached to the smooth surface of the foot layer; note the main body of the grains (foot layer) does not fill out the whole space of the reticulum (f). Specimens, TV44-S148218 (a–c), TV142-S170216 (d–f). Scale bars 300 Μm (a), 6 Μm (b, d–f), 3 Μm (c). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 42. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis sp. 1 (a–c) from an isolated anther and Piercipollis sp. 2 (d–f) from a coprolite (not shown); Torres Vedras locality, Portugal. a) Narrow elongate anther that yielded the pollen in (b) and (c); b) Distal view of pollen grain showing well-developed reticulum that is only loosely attached to the foot layer; note the smooth foot layer (arrowhead) of a grain from which the reticulum has become detached; c) Reticulum showing the smooth muri and long columellae that are mostly detached from the foot layer; d–f) Pollen grains in distal (d, e), and lateral views (f) showing the very long colpus and the well-developed reticulum only loosely attached to the smooth surface of the foot layer; note the main body of the grains (foot layer) does not fill out the whole space of the reticulum (f). Specimens, TV44-S148218 (a–c), TV142-S170216 (d–f). Scale bars 300 Μm (a), 6 Μm (b, d–f), 3 Μm (c).

Text-fig. 40. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartiopsis parva gen. et sp. nov. from an anther; Torres Vedras locality, Portugal. a) Holotype; elongated anther that yielded the pollen in this Text-figure; b–e) Pollen grains showing dense reticulate tectum supported by short columellae that are only loosely attached to the smooth surface of the foot layer: note dentate orbicules on inner surface of anther wall (b, arrowhead); f) Reticulate tectum showing smooth muri with small adhering orbicules (arrowhead); g) Pollen grains showing one in which the reticulum has been lost exposing the smooth surface of the foot layer; h) Fractured pollen wall showing reticulum, short columellae and thick foot layer; note granules on the inner surface of the apertural region (arrow). Specimen, TV39-S170217 (holotype). Scale bars 300 Μm (a), 6 Μm (b–e, g), 3 Μm (f, h). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 40. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartiopsis parva gen. et sp. nov. from an anther; Torres Vedras locality, Portugal. a) Holotype; elongated anther that yielded the pollen in this Text-figure; b–e) Pollen grains showing dense reticulate tectum supported by short columellae that are only loosely attached to the smooth surface of the foot layer: note dentate orbicules on inner surface of anther wall (b, arrowhead); f) Reticulate tectum showing smooth muri with small adhering orbicules (arrowhead); g) Pollen grains showing one in which the reticulum has been lost exposing the smooth surface of the foot layer; h) Fractured pollen wall showing reticulum, short columellae and thick foot layer; note granules on the inner surface of the apertural region (arrow). Specimen, TV39-S170217 (holotype). Scale bars 300 Μm (a), 6 Μm (b–e, g), 3 Μm (f, h).

Text-fig. 41. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis simplex gen. et sp. nov. from a pollen clump; Torres Vedras locality, Portugal. a) Holotype; pollen clump that yielded the pollen in this Text-figure; b–d) Pollen grains in distal (b), proximal (c) and lateral views (d) showing the very long colpus and the well-developed reticulum that is only loosely attached to the smooth surface of the foot layer; note the large angular lumina, smooth muri and short, sparsely scattered, columellae; note also that main body of pollen grains (foot layer) does not fill out the whole space of the reticulum; e) Reticulum showing the smooth muri loosely attached to the smooth surface of the foot layer by short, sparsely scattered, columellae. Specimen, TV44-S136755 (holotype). Scale bars 300 Μm (a), 6 Μm (b–d), 3 Μm (e). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 41. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis simplex gen. et sp. nov. from a pollen clump; Torres Vedras locality, Portugal. a) Holotype; pollen clump that yielded the pollen in this Text-figure; b–d) Pollen grains in distal (b), proximal (c) and lateral views (d) showing the very long colpus and the well-developed reticulum that is only loosely attached to the smooth surface of the foot layer; note the large angular lumina, smooth muri and short, sparsely scattered, columellae; note also that main body of pollen grains (foot layer) does not fill out the whole space of the reticulum; e) Reticulum showing the smooth muri loosely attached to the smooth surface of the foot layer by short, sparsely scattered, columellae. Specimen, TV44-S136755 (holotype). Scale bars 300 Μm (a), 6 Μm (b–d), 3 Μm (e).

Text-fig. 44. Scanning electron microscope (SEM) images of monocolpate pollen of Juhaszia portugallica gen. et sp. nov. from a fragment of a pollen sac (a–e) and stamen fragment and pollen of Dictyozonia pusilla gen. et sp. nov. (f–k); Torres Vedras locality, Portugal. a) Holotype; fragment of pollen sac that yielded the pollen in (b–e); b) Proximal surface of pollen grain showing reticulate sculpture in the equatorial regions grading into more a continuous foveolate tectum at the proximal pole; c) Reticulum on the proximal surface showing smooth muri delimiting large and small luminae. Muri supported by numerous short columellae that are only loosely attached to the smooth surface of the foot layer; d) Distal surface of pollen grain showing the long colpus and reticulate sculpture with a distinctive pattern of large and small luminae; e) Pollen grain from which the reticulum has become in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 44. Scanning electron microscope (SEM) images of monocolpate pollen of Juhaszia portugallica gen. et sp. nov. from a fragment of a pollen sac (a–e) and stamen fragment and pollen of Dictyozonia pusilla gen. et sp. nov. (f–k); Torres Vedras locality, Portugal. a) Holotype; fragment of pollen sac that yielded the pollen in (b–e); b) Proximal surface of pollen grain showing reticulate sculpture in the equatorial regions grading into more a continuous foveolate tectum at the proximal pole; c) Reticulum on the proximal surface showing smooth muri delimiting large and small luminae. Muri supported by numerous short columellae that are only loosely attached to the smooth surface of the foot layer; d) Distal surface of pollen grain showing the long colpus and reticulate sculpture with a distinctive pattern of large and small luminae; e) Pollen grain from which the reticulum has become

Text-fig. 49. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of Vedresia elliptica gen. et sp. nov. (a–c) and a multicarpellate fruit of Choffaticarpus compactus gen. et sp. nov. and associated pollen (d–j); Torres Vedras locality, Portugal. a) Holotype; small fruit showing apical stigmatic region with attached pollen; b) Group of pollen grains from the stigmatic surface of the fruit in (a) showing the long colpus and reticulate tectum; c) Pollen grains from the stigmatic surface showing the long colpus and well developed reticulum, with smooth muri and short columellae in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 49. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of Vedresia elliptica gen. et sp. nov. (a–c) and a multicarpellate fruit of Choffaticarpus compactus gen. et sp. nov. and associated pollen (d–j); Torres Vedras locality, Portugal. a) Holotype; small fruit showing apical stigmatic region with attached pollen; b) Group of pollen grains from the stigmatic surface of the fruit in (a) showing the long colpus and reticulate tectum; c) Pollen grains from the stigmatic surface showing the long colpus and well developed reticulum, with smooth muri and short columellae

Text-fig. 36. Scanning electron microscope (SEM) images of flower of Eckhartianthus lusitanicus gen. et sp. nov.; Torres Vedras locality, Portugal. a) Holotype; oblique lateral view of hypogynous flower showing perianth surrounding the stamens and protruding gynoecium; b–d) Pollen from the flower in (a) showing single distal aperture and reticulate exine with thick muri, small lumina and short, densely spaced columellae; e) Reticulum showing thick, smooth muri supported by numerous short columellae firmly attached to the smooth surface of the foot layer. Specimen, TV39-S101306 (holotype). Scale bars 300 Μm (a), 6 Μm (b–d), 3 Μm (e). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 36. Scanning electron microscope (SEM) images of flower of Eckhartianthus lusitanicus gen. et sp. nov.; Torres Vedras locality, Portugal. a) Holotype; oblique lateral view of hypogynous flower showing perianth surrounding the stamens and protruding gynoecium; b–d) Pollen from the flower in (a) showing single distal aperture and reticulate exine with thick muri, small lumina and short, densely spaced columellae; e) Reticulum showing thick, smooth muri supported by numerous short columellae firmly attached to the smooth surface of the foot layer. Specimen, TV39-S101306 (holotype). Scale bars 300 Μm (a), 6 Μm (b–d), 3 Μm (e).

Text-fig. 37. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia brevicolumella gen. et sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther partially split along the elongated sporangia; b) Dithecate, tetrasporangiate anther; c) Holotype; distal view of pollen grain from holotype showing the long colpus with a distinct margin and a heterobrochate reticulum; d) Distal view of pollen grain from a stamen fragment showing long colpus, colpus margin and heterobrochate reticulum; e) Inner surface of anther wall from stamen fragment showing densely spaced dentate orbicules; f) Lateral view of pollen grains from stamen showing the psilate to foveolate-microreticulate colpus margin; g) Detail of pollen grain from holotype showing the slightly rounded profile of the muri, the short, densely spaced columellae and the dentate orbicules. Specimens, TV44-S136662 (a), TV44-S149204 (b), TV44-S136760 (holotype; c, g), TV44-S148021 (d, e), TV44-S136752 (f). Scale bars 300 Μm (a, b), 6 Μm (c, d, f), 3 Μm (e, g). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 37. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia brevicolumella gen. et sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther partially split along the elongated sporangia; b) Dithecate, tetrasporangiate anther; c) Holotype; distal view of pollen grain from holotype showing the long colpus with a distinct margin and a heterobrochate reticulum; d) Distal view of pollen grain from a stamen fragment showing long colpus, colpus margin and heterobrochate reticulum; e) Inner surface of anther wall from stamen fragment showing densely spaced dentate orbicules; f) Lateral view of pollen grains from stamen showing the psilate to foveolate-microreticulate colpus margin; g) Detail of pollen grain from holotype showing the slightly rounded profile of the muri, the short, densely spaced columellae and the dentate orbicules. Specimens, TV44-S136662 (a), TV44-S149204 (b), TV44-S136760 (holotype; c, g), TV44-S148021 (d, e), TV44-S136752 (f). Scale bars 300 Μm (a, b), 6 Μm (c, d, f), 3 Μm (e, g).

Text-fig. 35. Scanning electron microscope (SEM) images of a fruit of Pennicarpus sp. and monocolpate pollen of Pennipollis sp. from a pollen clump; Torres Vedras locality, Portugal. a) Fruit of Pennicarpus sp. showing the poorly preserved fruit wall enclosing a single seed; b) Pollen clump that yielded the pollen in this Text-figure (c, d); c, d) Pollen grains showing the loose reticulum bearing small spinules that are often in two rows; note the smooth surface of the foot layer, the absence of columellae, the continuous muri bordering the apertures and spiny orbicules (arrowheads). Specimens, TV38-S174609 (a), TV44-S148015 (b–d). Scale bars 300 Μm (a, b), 6 Μm (c, d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 35. Scanning electron microscope (SEM) images of a fruit of Pennicarpus sp. and monocolpate pollen of Pennipollis sp. from a pollen clump; Torres Vedras locality, Portugal. a) Fruit of Pennicarpus sp. showing the poorly preserved fruit wall enclosing a single seed; b) Pollen clump that yielded the pollen in this Text-figure (c, d); c, d) Pollen grains showing the loose reticulum bearing small spinules that are often in two rows; note the smooth surface of the foot layer, the absence of columellae, the continuous muri bordering the apertures and spiny orbicules (arrowheads). Specimens, TV38-S174609 (a), TV44-S148015 (b–d). Scale bars 300 Μm (a, b), 6 Μm (c, d).