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Figure 193 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 193: Histogram showing maximized morphometric ratios of species (male) in genera Hoffmannius, Kochius, and Thorellius. See Figure 192 for data on female and Table 6 for additional data. Variance analysis p-values depicted in rectangles, Hoffmannius as compared to Thorellius and Kochius on left, and Kochius as compared to Thorellius on right. Statistically significant p-values (i.e., very small) indicated with black arrow. See Fig. 1 for explanation of histogram components.
Figure 192 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 192: Histogram showing maximized morphometric ratios of species (female) in genera Hoffmannius, Kochius, and Thorellius. See Figure 193 for data on male, and Table 6 for additional data. Variance analysis p-values depicted in rectangles, Hoffmannius as compared to Thorellius and Kochius on left, and Kochius as compared to Thorellius on right. Statistically significant p-values (i.e., very small) indicated with black arrow. See Fig. 1 for explanation of histogram components.
Figure 194 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 194: Morphometric ratios contrasting metasomal segments (length/width) for species in tribe Syntropini (male). Variance analysis p-values depicted in rectangles, Hoffmannius as compared to Kochius. Statistically significant p-values (i.e., very small) indicated with black arrow. See Fig. 1 for explanation of histogram components.
Figures 209–217 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 209–217: Kochius kovariki, sp. nov., female holotype. 209. Leg III tarsus, ventral view, showing median spinule row and four pairs of distal spinules. 210. Metasomal segment V, ventral view, showing narrowly bifurcated ventromedian carina. 211. Chelal movable fingers showing dentition pattern and position of trichobothria ib–it, fixed finger left and movable finger right. 212. Carapace with close-up of lateral eyes. 213. Stigma. 214. Chelicerae, ventral, external, and dorsal views. 215. Pecten. 216. Metasomal segments IV–V and telson, lateral view. 217. Telson, ventral view.
Figures 160–169 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 160–169: Metasomal segment IV, lateral and dorsal views, showing development of the terminus of the dorsal (D) and dorsolateral (DL) carinae for the three subfamilies of Vaejovidae. Subfamily Smeringurinae: 160. Paruroctonus stahnkei, male, Mesa, Arizona, USA. 161. Smeringurus aridus, male, ABDSP, California, USA. 162. Vejovoidus longiunguis, male, Las Bombas, Baja California Sur, Mexico. 163. Paravaejovis pumilis, male, Ciudad Constitución, Baja California Sur, Mexico. Subfamily Syntropinae: 164. Stahnkeus subtilimanus, ABDSP, California, USA. 165. Kochius punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 166. Hoffmannius viscainensis, male, Las Bombas, Baja California Sur, Mexico. 167. Hoffmannius gravicaudus, female, Santa Rosalia, Baja California Sur, Mexico. Subfamily Vaejovinae: 168. Vaejovis solegladi, female, Cuicatlan, Oaxaca, Mexico. 169. Pseudouroctonus reddelli, male, Comal Co., Texas, USA. AC = articulation condyle, td1 = terminal denticle of dorsal carina, td2 = terminal denticle of dorsolateral carina. After Soleglad & Fet (2003b), in part.
Figures 170–181 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 170–181: Telson, lateral view, of representative Syntropini species. 170. Kochius hirsuticauda, female, Indian Gorge Canyon, ABDSP, California, USA. 171. K. bruneus loretoensis, male, Loreto, Baja California Sur, Mexico. 172. K. punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 173. K. punctipalpi punctipalpi, male, Los Aripes, Baja California Sur, Mexico. Note the significant sexual dimorphism in this species. 174. K. cazieri, male, Cuatro Ciénegas, Coahuila, Mexico. 175. K. russelli female, Deming, New Mexico, USA. 176. Thorellius atrox, female, Colima, Colima, Mexico. 177. T. cristimanus, male, Autlán, Jalisco, Mexico. 178. Hoffmannius globosus, female, Zacatecas, Zacatecas, Mexico. 179. H. eusthenura, male, Cabo San Lucas, Baja California Sur, Mexico. 180. Syntropis williamsi, female holotype, Los Aripes, Baja California Sur, Mexico. 181. S. macrura, female, Isla Carmen, Baja California Sur, Mexico. (Figs. 180, 181 after Soleglad et al., 2007, in part).
Figures 182–190 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 182–190: Base of telson vesicle, dorsal view, showing development of vesicular "tabs" (pointed to by arrow) in subfamilies Syntropinae and Smeringurinae. Note the reduced tabs, the sharp granules essentially obsolete, in Smeringurus, Paravaejovis, and Vejovoidus, whereas in Paruroctonus and subfamily Syntropinae, the spine is distinct and well developed. 182. Hoffmannius waeringi, male, Indian Gorge Canyon, ABDSP, California, USA. 183. Kochius punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 184. Stahnkeus subtilimanus, female, Split Mountain, ABDSP, California, USA. 185. Paruroctonus becki, male, Cottonwood Springs, Joshua Tree National Monument, California, USA. 186. Paruroctonus silvestrii, Chihuahua Road, ABDSP, California, USA. 187. Paruroctonus luteolus, male, Palo Verde Wash, ABDSP, California, USA. 188. Smeringurus mesaensis, female, Palo Verde Wash, ABDSP, California, USA. 189. Paravaejovis pumilis, male, Ciudad Constitución, Baja California Sur, Mexico. 190. Vejovoidus longiunguis, female, Las Bombas, Baja California Sur, Mexico.
Figures 152–159 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 152–159: Diagrammatic view of chelal palm, proximal aspect, showing digital carinae (D1–D5). In particular, this perspective shows the profiled development of the subdigital (D2) carina and its relative position with respect to D1 and D3. See Figs. 140–145 for lateral view of D2 carina showing its relative length as compared to chelal palm length. 152. Kochius punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 153. K. bruneus loretoensis, male, Loreto, Baja California Sur, Mexico. 154. K. hirsuticauda, female, Indian Gorge Canyon, ABDSP, California, USA. 155. K. cazieri, male, Cuatro Ciénegas, Coahuila, Mexico. 156. K. russelli, female, Deming, New Mexico, USA. 157. Thorellius intrepidus, male, Mexico. 158. T. cristimanus, male, Autlán, Jalisco, Mexico. 159. T. atrox, female, Colima, Colima, Mexico. D1 = digital carina, D2 = subdigital carina, D3 = dorsosecondary carina, D4 = dorsomarginal carina, D5 = dorsointernal carina.
Figures 140–151 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 140–151: Chela, external view, of representative Kochius, Thorellius, Hoffmannius, and Syntropis species, showing carination and trichobothria pattern. In particular note the well developed subdigital (D2) carina, the mid-palm position of trichobothrium Dt, and location of Db dorsal of the digital (D1) carina. 140. Kochius punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 141. K. hirsuticauda, female, Indian Gorge Canyon, ABDSP, California, USA. 142. K. bruneus loretoensis, male, Loreto, Baja California Sur, Mexico. 143. K russelli, female, Deming, New Mexico, USA. 144. Thorellius intrepidus, male, Mexico. 145. T. atrox, female, Colima, Colima, Mexico. 146. Hoffmannius eusthenura, male, Cabo San Lucas, Baja California Sur, Mexico. 147. H. gravicaudus, female, Santa Rosalia, Baja California Sur, Mexico. 148. H. globosus, female, Zacatecas, Zacatecas, Mexico. 149. H. puritanus, male, Jasper Trail, ABDSP, California, USA. 150. H. viscainensis, female, Las Bombas, Baja California Sur, Mexico. 151. Syntropis williamsi, male paratype, Los Aripes, Baja California Sur, Mexico (after Soleglad et al., 2007: fig. 13, in part).
Figure 191 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 191: Metasomal segment (I–IV) proportions for subfamily Smeringurinae (ratios = segment width / segment length). Data include both adult males and females combined. These data show that Smeringurus segments are significantly thinner than in the other three genera, not only showing complete separation between standard error ranges, but for all data points. Statistical difference between Smeringurus and Paruroctonus is indicated by significant standard error separation and the negligible variance analysis value (p-value). In addition, segment III is at least twice as long as wide (indicated by 0.500 triangle marker) and segment IV is in general three times as long as wide (indicated by 0.333 triangle marker). The other genera cluster together, with Vejovoidus and Paravaejovis exhibiting the most stocky metasoma. The data, which encompass all species currently defined in Smeringurinae, were gathered from measurements calculated in this study as well as published measurements, primarily from those sources where the species were originally described (i.e., most measurements are from type specimens). See Fig. 1 for explanation of histogram components.
Figure 127 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 127: Method of measurements for carapace. Cara_L = carapace length; Cara_MW = carapace median width; MET_W = median eye tubercle width; MET_P = median eye tubercle position; and MED = median eye diameter.
Figures 57–66 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 57–66: Right hemispermatophore, dorsal view, of genera of subfamily Smeringurinae. Dotted line indicates position of ventral trough. Tribe Paravaejovini: 57. Paravaejovis pumilis, Ciudad Constitución, Baja California Sur, Mexico. Tribe Smeringurini: 58. Paruroctonus boreus, Mercury, Nevada, USA, dorsal and dorsoexternal views. 59. Paruroctonus gracilior, Lajitas, Brewster Co., Texas, USA. 60. Paruroctonus silvestrii, Chihuahua Road, ABDSP, California, USA. 61. Paruroctonus utahensis, Bluff, Utah, USA. 62. Smeringurus mesaensis, Borrego Springs, California, USA, dorsal and internoventral views. 63. Smeringurus aridus, Palo Verde Wash, ABDSP, California, USA. 64. Smeringurus vachoni immanis, 1000 Palms, California, USA. 65. Smeringurus grandis, Oakies Landing, Baja California, Mexico. 66. Vejovoidus longiunguis, Las Bombas, Baja California Sur, Mexico.
Figures 41–43 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 41–43: Right hemispermatophore of Thorellius cristimanus, Autlán, Jalisco, Mexico. 41. Dorsal view; dotted line indicates position of ventral trough, see Fig. 43. 42. Internoventral view showing partial view of embedded mating plug. 43. Ventral view showing embedded mating plug partially visible on distal aspect, barb on left pointing inwards into capsular area. See Fig. 75 for extracted mating plug, dorsal view (barb pointing outward), showing toothed (16 teeth) distal barb.
Figures 128–139 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 128–139: Carapace, anterior half, of representative Syntropini species, emphasizing differences in the anterior emargination and median indentation. 128. K. punctipalpi punctipalpi, female, Cabo San Lucas, Baja California Sur, Mexico. 129. K. bruneus loretoensis, male, Loreto, Baja California Sur, Mexico. 130. K. hirsuticauda, female, Indian Gorge Canyon, ABDSP, California, USA. 131. K. cazieri, male, Cuatro Ciénegas, Coahuila, Mexico. 132. K russelli, female, Deming, New Mexico, USA. 133. K. kovariki, female holotype, Durango, Durango, Mexico. 134. T. atrox, female, Colima, Colima, Mexico. 135. T. cristimanus, male, Autlán, Jalisco, Mexico. 136. H. eusthenura, male, Cabo San Lucas, Baja California Sur, Mexico. 137. H. gravicaudus, female, Santa Rosalia, Baja California Sur, Mexico. 138. H. globosus, female, Zacatecas, Zacatecas, Mexico. 139. Syntropis aalbui, holotype female, Cataviña, Baja California, Mexico (after Soleglad et al., 2007: fig. 17, in part).
Figures 103–113 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 103–113: Leg tarsus, ventral view, of select Syntropinae species. Tribe Syntropini, exhibiting multiple distal spinule pairs. 103. Kochius punctipalpi punctipalpi, female, right leg III. 104. Thorellius cristimanus, male, right leg IV. 105. Hoffmannius eusthenura, male, right leg III. 106. Syntropis williamsi, female holotype, right leg IV (after Soleglad et al., 2007: fig. 8, in part). 107. Kochius cazieri, male, right leg III. 108. Thorellius atrox, female, right leg III. 109. Hoffmannius gravicaudus, female, right leg III. 110. Syntropis aalbui, female holotype, leg IV (after Soleglad et al., 2007: fig. 20, in part). Tribe Stahnkeini, exhibiting a single distal spinule pair. 111. Serradigitus wupatkiensis, female, right leg III. 112. Serradigitus adcocki, female, right leg III. 113. Stahnkeus subtilimanus, female, right leg III.
Figure 13 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 13: Trichobothrial pattern of Pseudouroctonus williamsi, male, Santa Ysabel Reserve, San Diego Co., California, USA.
Figures 23–26 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 23–26: Trichobothrial pattern of pedipalp patella, external view, of genus Thorellius showing orthobothriotaxy. 23. T. intrepidus, male, Mexico. 24. T. cristimanus, male, Autlán, Jalisco, Mexico. 25. T. atrox, female, Colima, Colima, Mexico. 26. T. occidentalis, female, Acapulco, Guerrero, Mexico. Abbreviations of trichobothria: eb = external basal; esb = external suprabasal; em = external median; est = external subterminal; et = external terminal; v3 = ventral trichobothrium number 3.
Figure 7 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 7: Trichobothrial pattern of Paravaejovis pumilis, male, Ciudad Constitución, Baja California Sur, Mexico.
Figure 4 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figure 4: Trichobothrial pattern of Paruroctonus surensis, male, Las Bombas, Baja California Sur, Mexico.
Figures 75–88 in Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae)
Figures 75–88: Representative hemispermatophore mating plugs of subfamilies Syntropinae, Smeringurinae, and Vaejovinae. Tribe Syntropini, note toothed terminus. 75. Thorellius cristimanus, Autlán, Jalisco, Mexico. 76. Kochius punctipalpi punctipalpi, Los Aripes, Baja California Sur, Mexico. 77. Kochius hirsuticauda, Indian Gorge Canyon, ABDSP, California, USA. 78. Kochius cazieri, Cuatro Ciénegas, Coahuila, Mexico. 79. Hoffmannius eusthenura, Cabo San Lucas, Baja California Sur, Mexico. 80. Hoffmannius puritanus, Jasper Trail, ABDSP, California, USA. 81. Hoffmannius confusus, Mesa, Arizona, USA. 82. Syntropis macrura (after Stockwell, 1989: fig. 228, in part). Tribe Stahnkeini, note smooth terminus. 83. Serradigitus wupatkiensis, ventral view, Wupatki Ruins, Wupatki National Monument, Arizona, USA. 84. Stahnkeus subtilimanus, ventral view, Vallecito Creek, Carrizo Badlands, ABDSP, California, USA. Tribe Smeringurini, note smooth terminus. 85. Paruroctonus silvestrii, Chihuahua Road, ABDSP, California, USA. 86. Paruroctonus boreus, Mercury, Nevada, USA. Subfamily Vaejovinae, note smooth terminus. 87. Pseudouroctonus minimus castaneus, Vista, San Diego Co., California, USA. 88. Pseudouroctonus williamsi, Santa Ysabel Reserve, San Diego Co., California, USA.
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