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Figure 2 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 2 Imagines of the C. cyprius-group in lateral view, proboscis retracted (C. yemenensis female dbM 4944 male dbM 4945, C. osseus female dbM 4617 male dbM 5521, C. ivanovi sp. nov. female dbM 5493 male dbM 5498, C. cagrii sp. nov. female dbM 5607 male dbM 5619, C. clarus female dbM 5674).

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Figure 11 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 11 Microphotos of male genitalia in dorsal view aC. rugiceps (dbM 5450) bC. cyprius (dbM 5439) cC. yemenensis (dbM 5532) dC. osseus (dbM 5521) eC. ivanovi sp. nov. (dbM 5498) fC. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).

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Figure 10 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 10 Male genitalia in dorsal view aC. rugiceps (dbM 2834) bC. cyprius (dbM 5452) cC. yemenensis (dbM 5531) dC. osseus (dbM 5521) eC. ivanovi sp. nov. (dbM 5497) fC. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).

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Figure 6 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 6 a, b Metasomal tergum II and III of female in dorsal view aC. rugiceps (dbM 5446) bC. ivanovi sp. nov. (dbM 5481) c–f mesonotum of female in dorsal view cC. ivanovi sp. nov. (dbM 5492) dC. osseus (dbM 4617) eC. yemenensis (dbM 5528) fC. clarus (dbM 4616).

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Figure 15 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 15 Female antennal articles A1–A6 in medial view (scaled to same size to show different proportions of A3) aC. clarus (dbM 5674) bC. osseus (dbM 5678).

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Figure 16 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 16 Neighbour joining BOLD taxon ID tree of 18 specimens of the C. cyprius-group based on COI-5P gene sequences (individually marked by BOLD process ID and assigned to species a priori by morphological characters; COI-5P gene sequence of C. osseus and C. clarus unknown).

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Figure 17 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 17 Geographic distribution of members of the C. cyprius-group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www.naturalearthdata.com).

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Figure 18 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 18 a–e, g, h Flower visiting behaviour of Celonites ivanovi sp. nov. females at flowers of Heliotropium styligerum and behaviours associated with it (details see text) f habitat of C. ivanovi sp. nov. at locality 2 in the vicinity of Maydanskoye, Dagestan i habitat of C. cagrii sp. nov. near Akşar, Turkey (aspect in autumn).

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Figure 5 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 5 a, b Female clypeus in frontal view aC. cyprius (dbM 5431) bC. ivanovi sp. nov. (dbM 5490) c, d female mandible outside cC. cyprius (dbM 5431) dC. ivanovi sp. nov. (dbM 5490) e, f female ocelli and vertex in dorsal view eC. ivanovi sp. nov. (dbM 5492) fC. cagrii sp. nov. (dbM 5609).

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Figure 14 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 14 Male fused sterna VII+VIII in ventral view aC. rugiceps (dbM 5289) bC. cyprius (dbM 5439) cC. yemenensis (dbM 5532) dC. osseus (dbM 5521) eC. ivanovi sp. nov. (dbM 5498) fC. cagrii sp. nov. (dbM 5619).

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Figure 4 from: Mauss V, Fateryga AV, Yildirim E, Carpenter JM (2022) Contribution to the taxonomy, bionomics and distribution of the Palaearctic Celonites cyprius-group (Hymenoptera, Vespidae, Masarinae) with the description of two new species from the North Caucasus and East Anatolia. Journal of Hymenoptera Research 89: 109-155. https://doi.org/10.3897/jhr.89.79832

Figure 4 Female head in frontal view aC. yemenensis (dbM 5528) bC. clarus (dbM 4616) cC. cyprius (dbM 5431) dC. osseus (dbM 5678) eC. ivanovi sp. nov. (dbM 5490) fC. cagrii sp. nov. (dbM 5609).

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Figures 9-14 from: Mauss V, Müller A, Prosi R (2018) Flower associations and nesting of the pollen wasp Quartinia major Kohl, 1898 (Hymenoptera, Vespidae, Masarinae) in Morocco. Journal of Hymenoptera Research 62: 15-31. https://doi.org/10.3897/jhr.62.22879

Figures 9-14 Flower visiting behaviour of females of Quartinia major. 9 Nectar uptake from disc florets on capitulum of Asteriscus graveolens 10 Female with protruded proboscis taking up nectar from disc florets of Pulicaria mauritanica 11 Ingestion with the mouthparts of pollen grains that had accumulated on the fore legs 12 Female pressing pollen out of the corolla tube of a disc floret of P. mauritanica with her mouthparts, while the pollen grains accumulate above the corolla in front of her head surrounded by her antennae 13 Pollen uptake from disc floret of P. mauritanica a Female pressing pollen out of the corolla tube b female immediately afterwards ingesting pollen supported by her fore legs 14 Female brushing pollen from her exoskeleton with rapid alternating movements of her fore legs.

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Figures 3-8 from: Mauss V, Müller A, Prosi R (2018) Flower associations and nesting of the pollen wasp Quartinia major Kohl, 1898 (Hymenoptera, Vespidae, Masarinae) in Morocco. Journal of Hymenoptera Research 62: 15-31. https://doi.org/10.3897/jhr.62.22879

Figures 3-8 3 Female of Quartinia major visiting disc florets on a capitulum of Pulicaria mauritanica. The proterandric disc florets open from the outer ones inwards. The mouthparts of the female are situated in the zone where disc florets are in an early male phase of anthesis 4 Location of nest CH of Q. major at locality I 5 Nest entrance of nest GB (viewed at an angle) 6 Nest entrance of nest CH (viewed from above) 7 Female of nest CH partly backed out of the turret moving slightly around her longitudinal axis with her mouthparts orientated towards the inner surface of the turret wall 8 Female of nest CH backing out of the nest carrying a load of soil particles with her mouthparts (visible between fore and mid femur).

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Figure 15 from: Mauss V, Müller A, Prosi R (2018) Flower associations and nesting of the pollen wasp Quartinia major Kohl, 1898 (Hymenoptera, Vespidae, Masarinae) in Morocco. Journal of Hymenoptera Research 62: 15-31. https://doi.org/10.3897/jhr.62.22879

Figure 15 Schematic vertical cross sections of the nest architecture and brood cell arrangement of nest CH and nest GB of Quartinia major investigated on 15 April, 2017 at locality I (c = brood cell with cell number, ls = little stone, s = main shaft, t = turret).

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Figures 1-2 from: Mauss V, Müller A, Prosi R (2018) Flower associations and nesting of the pollen wasp Quartinia major Kohl, 1898 (Hymenoptera, Vespidae, Masarinae) in Morocco. Journal of Hymenoptera Research 62: 15-31. https://doi.org/10.3897/jhr.62.22879

Figures 1-2 Habitat of Quartinia major. 1 Locality I, ruderal road side 20 km to the north of Tafraout, with yellow flowering dwarf shrubs of Pulicaria mauritanica. Two nests were found close to the little white flag in the foreground on the left 2 Locality II, wadi 2.75 km to the south-west of Ifrane Atlas Saghir, with yellow flowering plants of Asteriscus graveolens that were visited by females of Q. major.

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Figures 7-9 from: Mauss V, Prosi R (2018) Identity and distribution of Celonites hermon Gusenleitner, 2002 (Hymenoptera, Vespidae, Masarinae) from the Middle East with a description of the hitherto unknown male. Journal of Hymenoptera Research 66: 55-70. https://doi.org/10.3897/jhr.66.29795

Figures 7-9 Head of male Celoniteshermon in dorsal view (dbM No. 3456) 8 Metasomal sternum VIII of male of C.hermon in ventral view (dbM No. 3689) 9 Male genital of Celoniteshermon (dbM No. 3689) a in dorsal b ventral view.

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Figures 2-6 from: Mauss V, Prosi R (2018) Identity and distribution of Celonites hermon Gusenleitner, 2002 (Hymenoptera, Vespidae, Masarinae) from the Middle East with a description of the hitherto unknown male. Journal of Hymenoptera Research 66: 55-70. https://doi.org/10.3897/jhr.66.29795

Figures 2-6 Head of female of Celoniteshermon (dbM No. 3685) 2 in frontal view 3 in lateral view showing pollen collecting apparatus with "knobbed" setae on frons and clypeus 4 in dorsal view 5–6 Metasoma of Celoniteshermon in dorsal view 5 female (dbM No. 3685) 6 male (dbM No. 3689).

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Figure 11 from: Mauss V, Prosi R (2018) Identity and distribution of Celonites hermon Gusenleitner, 2002 (Hymenoptera, Vespidae, Masarinae) from the Middle East with a description of the hitherto unknown male. Journal of Hymenoptera Research 66: 55-70. https://doi.org/10.3897/jhr.66.29795

Figure 11 Geographic distribution of the species of the Celonitesabbreviatus-complex occurring in the Middle East. Records of Celonitesandreasmuelleri taken from Mauss 2013 and Mauss unpublished, records of Celonitestauricus according to Mauss et al. 2016 and Mauss unpublished.

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Figure 10 from: Mauss V, Prosi R (2018) Identity and distribution of Celonites hermon Gusenleitner, 2002 (Hymenoptera, Vespidae, Masarinae) from the Middle East with a description of the hitherto unknown male. Journal of Hymenoptera Research 66: 55-70. https://doi.org/10.3897/jhr.66.29795

Figure 10 Male genital of Celoniteshermon (dbM No. 3689) in dorsal (left) and ventral view (right). Setae only shown on one side of each drawing. Nomenclature follows that of Birket-Smith (1981): ad, aedoeagus; at, apodema thyrsos; cu, cupula; dc, dorso-medial concavity of the posterior margin of the stipes; ha, harpide; mp, medial process of volsella; pp, posterior process of volsella; sl, shovel-like dorso-medial lobe of harpide; sp, stipes; ty, thyrsos; vo, volsella.

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Abb. 8-11 in Über Quartinia-Arten aus der Arabischen Halbinsel (Hymenoptera: Vespidae, Masarinae)

Abb. 8-11: (8) Quartinia emirata nov.sp. ♀, Habitus; (9) Quartinia emirata nov.sp. ♀, Stirn; (10) Quartinia emirata nov.sp. ♀, Tergite; (11) Quartinia emirata nov.sp. ♂, Clypeus;

opencc-by-4.0Dec 2018View details →

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