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Figure 2 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 2 - Map indicating geographic position of massifs and caves with Zospeum species. Dot in Triangle, Zospeum zaldivarae sp. n.: Cueva de Las Paúles (locus typicus); Dot in circle, Zospeum vasconicum sp. n.: Cueva de la Ermita de Sandaili (locus typicus); Simple orange circles, Zospeum vasconicum sp. n. localities: Cueva de Otxas; Cueva Silibranka-2; Cueva del Cranéo.

Figure 1 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 1 - Measurement on Zospeum shells in frontal view. Abbreviations: HLW height of last whorl PH peristome height PD peristome diameter SA spire angle SD shell diameter SH shell height.

Figure 4 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 4 - Zospeum vasconicum sp. n., A, F–I shells from Cueva Silibranka-2 in frontal view (RMNH.5003915) B–E different views of specimen A; H shell with window cut in body whorl exposing columella. Scale bar 0.5 mm.

Figure 7 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 7 - Zospeum zaldivarae sp. n., A–F different views of edentate paratype shell (RMNH.234152) D aperture in oblique view G paratype shell (UPV/EHU-FC: 72) with window in body whorl exposing columella.

Figure 3 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 3 - Zospeum vasconicum sp. n., A–E different views of holotype (MNCN 15.05/60147H) F–I paratype shells (MNCN 15.05/60147P) H paratype shell with window cut in body whorl exposing columella. Scale bar 0.5 mm.

Figure 6 from: Jochum A, de Winter AJ, Weigand AM, Gómez B, Prieto C (2015) Two new species of Zospeum Bourguignat, 1856 from the Basque-Cantabrian Mountains, Northern Spain (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 483: 81-96. https://doi.org/10.3897/zookeys.483.9167

Figure 6 - Zospeum zaldivarae sp. n., A, E–F different views of holotype shell (MNCN 15.05/60148H) D aperture in slightly oblique view showing apertural barriers.

Figure 8 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 8 - NanoCT−SEM Plate 2. A–F Zospeum isselianum, neotype (CSR SASA 37013), Turjeva jama G–H Zospeum isselianum (SMNH 2216), Jama na Zgornjih Brsnikih I–L Zospeum sp. (CSR SASA 37698a), Konečka zijalka.

Figure 2 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 2 - Historical material: A Zospeum isselianum syntype (MZUT M3232): upper mountains of Natisone River valley B–F Zospeum isselianum topotype: Turjeva jama G–P Specimens labelled as "Zospeum alpestre" in the Bourguignat Collection (MHNG 7898/2) from "Cav[erne]. de Carniole".

Figure 17 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 17 - Light micrograph of a section of the reproductive tract of Zospeum sp. (Konečka zijalka) (CSR SASA 21675). A Gonad (g) with acinus of the ovotestis including the germinal epithelial ring (ger) surrounding the acinar lumina (al), sustentacular cells (Sertoli cells) (sc) and oogonia (og) B Digestive gland (dg), hermaphroditic duct (hd) lined with cilia (c), posterior loop of the intestine (pi), receptaculum seminis (rs) containing allospermatozoa with spermatozoa oriented towards and apical ends touching the epithelial cells, and long motile cilia (ce) of intestine (i).

Figure 16 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 16 - Light micrograph of the histological appearance of the digestive tract of Zospeum sp. (Konečka zijalka) (CSR SASA 21675). A−B Bilobed muscular gizzard (mg) with epithelium showing columnar cells (cc) with microvillous apical surfaces (mv), posterior loop of the intestine (pi) with ciliated epithelium (ce) B Elaborate muscular plates (mp), thick cuticle (cu), highly ciliated rectum (r), gastric pouch (gp), digestive gland (dg) and hermaphroditic duct.

Figure 14 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 14 - Images of the radula of Zospeum isselianum (Turjeva jama) (CSR SASA 37013 conspecific) and Zospeum sp. (Konečka zijalka) (CSR SASA 21675). A−D SEM of the radular ribbon of topotypic Zospeum isselianum A Radular ribbon showing adhesive layer of contact zone with odontophore B SEM overview showing rachidian teeth (r) and lateral teeth (l) C Rows of lateral teeth bearing fine medial groove D Marginal teeth E Light micrograph showing the histological appearance of the radular sheath of Zospeum sp. (Konečka zijalka) showing the inferior epithelium (ie) attached to the radular membrane (rm) mineralizing cells (mc) with degenerating nuclei and radular teeth (rt).

Figure 4 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 4 - Overview of morphological assignments, molecular species delimitation results and integrative taxonomic treatment of each of the four clusters, which are indicated by the longitudinal black bars for either the ABDG or the SP approach. Numbers indicate individual specimens; ABGD = Automatic Barcode Gap Discovery; SP = Statistical Parsimony.

Figure 13 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 13 - A Light micrograph showing the histological appearance of Zospeum sp. (Konečka zijalka) (CSR SASA 21675) with the mantel cavity (mc), mucus glands (mg), oesophagus (oe) bearing thick cuticular sheath (cs) and leading into the crop (c), the buccal ganglia (bg), and the cephalopodial portion of the reproductive system including penis (p) and vas deferens (vd) B Paired cerebral ganglia united by the cerebral commissure situated just dorsal to the origin of the oesophagus in the buccal mass.

Figure 12 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 12 - Light micrograph showing histological overview of the digestive system of Zospeum sp. (Konečka zijalka) (CSR SASA 21675). A Mouth and buccal apparatus. Mouth slit between oral lappets (m), crescent-shaped jaw (j), radula (r), pharynx (ph), digestive gland (dg), intestine (i), kidney (k), columellar muscle (cm), and pedal ganglia (peg) B Mantle gland (mag), sigmoid intestine (si), radular sac (rs), cerebral ganglion (cg), mucus glands (mg), kidney (k), heart showing auricle and ventricle (h), digestive gland (dg), columellar muscle (cm), oesophagus (oe) and salivary glands (sg).

Figure 7 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 7 - Kimura−2-parameter (K2P) genetic distance table in %. Delineated species are marked with boxes. For each individual specimen, its morphospecies designation and genetic distance data to all remaining specimens are given.

Figure 11 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 11 - Type and shell images Plate 5: A–E Zospeum kupitzense holotype: (SMF 256354) F–H Zospeum kupitzense (SMNH 3291), Ložekarjeva zijalka I–M Zospeum amoenum (RS59), Potočka zijalka.

Figure 10 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 10 - NanoCT−SEM Plate 4. A–F Zospeum kupitzense (RS3291), Ložekarjeva zijalka G. Zospeum alpestre (SMNH 2216), Kamniška jama H, I−L Zospeum isselianum Ihanščica cave J−K Zospeum amoenum (RS59), Potočka zijalka.

Figure 1 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 1 - Zospeum isselianum potential distribution collated from the literature and museum collections (I. Sajko, CSR SASA).

Figure 15 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 15 - Light micrograph showing histological appearance of the radular complex of Zospeum sp. (Konečka zijalka) (CSR SASA 21675). A Odontoblasts (odb) grouped in lower posterior section of radular sheath, radular teeth (rt), odontophore (od), and collostyle (col) B Section through an acinus of the ovotestis showing some stages of development of sustentacular cells (Sertoli cells) (sc) with spermatogonia (spg), spermatids (sp) and oogonia (og).

Figure 3 from: Jochum A, Slapnik R, Klussmann-Kolb A, Páll-Gergely B, Kampschulte M, Martels G, Vrabec M, Nesselhauf C, Weigand AM (2015) Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeum isselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165. https://doi.org/10.3897/subtbiol.16.5758

Figure 3 - Geographical position and river drainage systems of the Julian Alps and the Kamnik Savinja Alps range. Sampling sites: 1 Turjeva jama 2 Ložekarjeva jama 3 Konečka zijalka 4 Potočka zijalka 5 Jama na Zgornjih Brsnikih 6 Tomažičeva zijalka 7 Kamniška jama 8 Ihanščica cave. Digital terrain model from Jarvis et al., 2008. River network data courtesy of Public Information of Slovenia, the Surveying and Mapping Authority of the Republic of Slovenia, DPK1000V (2008).