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FIG. 11. — Paralogania kachanovi n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 11. — Paralogania kachanovi n. sp., morphological set of the scales; A-E, head scales; F-H, transitional scales; I-T, trunk scales; L, holotype; A-I, J1, K, N, T1, scales in crown view; J2, L, M, O, Q, scales in lateral view; T2, scale in base view; P, R, scales in oblique postero-lateral view; S, scale in oblique lateral view; "×" shows the anterior part of the scale; A-K, LIG 35-039-LIG 35-049; L, LIG 35-050; M-P, LIG 35-051-LIG 35-054; Q-T, LIG 35-873-LIG 35-876; sample 12, Pioneer Island, Pridoli, Upper Silurian. Scale bar: 0.3 mm.
FIG. 7. — Paralogania menneri n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 7. — Paralogania menneri n. sp.; A, transitional scale; B-O, transitional(?) scales; P-T, trunk scales; A-C, E, F, H, J, L, S, scales in crown view; D, a scale in oblique lateral view (posterior end on the left); G, P, scales in base view; N, a scale in lateral view; I, magnified of H; K, magnified of J; M, magnified of L; O, magnified of N; R, magnified of P; T, magnified of S; A, Pi 7618; B, Pi 7607; C, D, Pi 7605; E, Pi 7604; F, LIG 35-871; G, LIG 35-872; H, I, LIG 35-877; J, K, holotype, LIG 35-878; L, M, LIG 35-879; N, O, LIG 35-880; P, R, LIG 35-881; S, T, LIG 35-882; A-E, sample MF 11-1, Matusevich River, October Revolution Island, Ust-Spokojnaya Formation, Ludlow; F, G, sample 18041-14, Komsomolets Island, Ludlow; H, J, L, N, P, S, sample 47-14, Spokojnaya River, October Revolution Island, Ust-Spokojnaya Formation, Ludlow, Upper Silurian. Scale bars: A, H-O, R, T, 0.1 mm; B-G, P, S, 0.2 mm.
FIG. 6 in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 6. — Microstructure of the scales; A-E, Paralogania ex gr. martinssoni (Gross, 1967); F, G, Thelodus sp. indet.; H, I, Lanarkia? sp.; A, Pi 6378; B, Pi 6362; C, Pi 6370; D, Pi 6368; E, Pi 6446; F, Pi 6377; G, Pi 6373; H, I, Pi 6371; sample 2055 zh, Pioneer Island, upper part of Ludlow, Upper Silurian. Abbreviations: b, base; cr, crown; dc, dentine canal; dt, dentine tubules; mpc, main pulp canal; n, neck; pcv, pulp cavity; wdt, wide dentine tubules. Scale bars: 0.1 mm.
FIG. 10. — Paralogania wilsoni n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 10. — Paralogania wilsoni n. sp., microstructure of the scales; A, Pi 7637; B, Pi 7638; C, Pi 7639; D, Pi 7640; E, F, Pi 7641; G, Pi 7642; H, Pi 7643; sample 157-2, Spokojnaya River, October Revolution Island, Krasnaya Bukhta Formation, Pridoli, Upper Silurian. Abbreviations: b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck; mpc, main pulp canal; pc, lateral pulp canal. Scale bars: 0.1 mm.
FIG. 20. — Thelodus matukhini n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 20. — Thelodus matukhini n. sp., morphological set of scales; A-C, head scales; D-G, transitional scales; H-S, trunk scales; T, pugniformis-type of the scale; A-C, D1, E-H, I1, J-O, P2, Q, R2, S, T1, scales in crown view; D2, scale in base view; I2, P1, R1, T2, scales in lateral view; LIG 35-281-LIG 35-300; sample 18041-14, Komsomolets Island, Ludlow, Upper Silurian. Scale bar: 0.3 mm.
FIG. 5 in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 5. — Set of scales of Paralogania ex gr. martinssoni (Gross, 1967); A, B, head scales; C-I, O, transitional scales; J-N, P-U, trunk scales; V-Z, fin scales; A, Pi 5605; B, Pi 5590; C, Pi 5607; D, Pi 5609; E, Pi 5611; F, Pi 5613; G, Pi 5617; H, Pi 5595; I, Pi 5591; J, Pi 5604; K, Pi 6772; L, Pi 5588; M, Pi 5606; N, Pi 5601; O, Pi 5600; P, Pi 6773; Q, Pi 6775; R, Pi 6776; S, Pi 6774; T, Pi 5612; U, Pi 5614; V, Pi 6779; W, Pi 5949; X, Pi 6777; Y, Pi 6778; Z, Pi 5948; sample 2055 zh, Pioneer Island, upper part of Ludlow, Upper Silurian (drawings after Märss 1986b: fig. 14). Scale bar: 0.5 mm.
FIG. 3. — A-M in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 3. — A-M, Paralogania ex gr. martinssoni (Gross, 1967); A, transitional scale; B-H, L, M, trunk scales; I, fin scale; J, K, scales of unknown position; A-E, G, I-L, scales in crown view; F, a scale in base view; H, M, scales in lateral view; N, Lanarkia? sp., a scale in lateral view; O-T, Phlebolepis elegans Pander, 1856; O-R, scales in crown view; S, magnified scale of Fig. 3Q showing the finely striated crown margin; T, scale in base view; O-Q, S, cephalo-pectoral scales; R, precaudal scale; T, precaudal? scale; A, Pi 7612; B, Pi 7609; C, Pi 7610; D, Pi 6845; E, Pi 7616; F, Pi 7617; G, H, Pi 7615; I, Pi 6847; J, Pi 6846; K, Pi 7608; L, LIG 35-900; M, LIG 35-901; N, LIG 35-902; O, Pi 7564; P, Pi 7560; Q, Pi 7562; R, Pi 7563; S, Pi 7562; T, Pi 7566; A-K, sample MF 11-1, Matusevich River, October Revolution Island, Ust-Spokojnaya Formation, Ludlow; L-N, sample 5m-1, Pioneer Island; O-T, sample MF 157-4, Spokojnaya River, October Revolution Island, Ludlow, Upper Silurian. Scale bars: A-D, F, G, K, S, 0.1 mm; E, H-J, M-O, Q, R, T, 0.2 mm; L, P, 0.3 mm.
FIG. 18 in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 18. — Set of scales of Thelodus visvaldi n. sp.; A-G, head scales; H-T, trunk scales; A, B1, C-G, H1, I1, J-M1, N-T, scales in crown view; B2, scale in base view; H2, I2, M2, scales in lateral view; LIG 35-120-LIG 35-139; sample 5m, Pioneer Island, Ludlow, Upper Silurian. Scale bar: 0.3 mm.
FIG. 17. — Thelodus visvaldi n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 17. — Thelodus visvaldi n. sp.; A, head scale; B, D-I, K-O, trunk scales; C, fin scale?; J, scale of unknown position; A-D, F-J, M-O, scales in crown view; E, N, scales in oblique lateral view; K, scale in lateral view; L, scale in base view; A, Pi 7541; B, Pi 7556; C, Pi 7549; D, E, Pi 7544; F, Pi 7542; G, LIG 35-903, the scale is lost; H, LIG 35-904; I, Pi 7545; J, Pi 7559; K, LIG 35-905; L, LIG 35- 906; M, Pi 7553; N, Pi 7554; O, Pi 7555; A-F, I, J, M-O, MF 34-1, Cape October, October Revolution Island; G, H, K, L, 5m-1, Pioneer Island; Ludlow, Upper Silurian. Scale bars: A-C, F-K, M-O, 0.1 mm; D, E, L, 0.2 mm.
FIG. 13. — Valiukia flabellata n. gen., n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 13. — Valiukia flabellata n. gen., n. sp.; A, B, head scales; C-H, transitional scales; I, trunk(?) scale; J, K, posterior part of trunk scales; A-E, G, I, scales in crown view; F, magnified part of E; H, magnified part of G; J, K, scales in base view; A-H, LIG 35-838-LIG 35-843; I-K, LIG 35-848-LIG 35-850; sample 2-23, Matusevich River, October Revolution Island, Ust-Spokojnaya Formation, Ludlow, Upper Silurian. Scale bars: 0.1 mm.
FIG. 2. — A-I, Thelodus matukhini n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 2. — A-I, Thelodus matukhini n. sp., scales in crown view; A, LIG 35-859; B, holotype, LIG 35-860; C, LIG 35-861; D, LIG 35- 862; E, Pi 7552; F, Pi 7550; G, Pi 7547; H, Pi 7551; I, Pi 7557; A-D, sample 18041-14, Komsomolets Island, Ludlow, Upper Silurian; E-I, sample MF 34-1, cape October (Octyabrskij), October Revolution Island, Ludlow, Upper Silurian; J-Q, Loganellia cuneata (Gross, 1947); J, L, O-Q, scales in crown view; K, M, scales in oblique lateral crown view; N, a scale in lateral view; J-Q, LIG 35-863- LIG 35-870; sample 2-23, Matusevich River, October Revolution Island, Ust-Spokojnaya Formation, Ludlow, Upper Silurian. Scale bars: A, C, E-H, J-P, 0.1 mm; B, D, I, Q, 0.2 mm.
FIG. 9. — Paralogania wilsoni n in Upper Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 9. — Paralogania wilsoni n. sp.; A, head scale; B-D, transitional scales; E-H, most probably the scales of leading edges of the fins; I-P, trunk scales; Q, a fin scale; A-D, F, G, I-Q, the scales in crown view; E, H, the scales in lateral view; A, Pi 7603; B, Pi 7594; C, Pi 7596; D, Pi 7601; E, Pi 7593; F, Pi 7591; G, Pi 7597; H, Pi 7602; I, Pi 7592; J, Pi 7599; K, Pi 7600; L, Pi 7595; M, holotype, Pi 7598; N, Pi 7590; O, Pi 7589; P, Pi 7587; Q, Pi 7588; sample MF 157-2, Spokojnaya River, October Revolution Island, Krasnaya Bukhta Formation, Pridoli, Upper Silurian. Scale bars: A-F, I, L, N-Q, 0.1 mm; G, H, J, K, M, 0.2 mm.
FIG. 15. — Thelodus calvus n in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 15. — Thelodus calvus n. sp., microstructure of the scales; A, Pi 7631; B, Pi 7632; C, D, Pi 7633; E, Pi 7634; F, Pi 7635; G, Pi 7636; sample MF 46-23, Ushakov River, October Revolution Island, Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine "pegs" in the base; b, base; cr, crown; dc, dentine canal; dt, dentine tubules; n, neck, pc, pulp canal; pcv, pulp cavity. Scale bars: 0.1 mm.
FIG. 11. — Paralogania consimilis n in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 11. — Paralogania consimilis n. sp.; A-C, head scales; D, G, transitional scales; E, F, H-T, trunk scales; A-D, F-M, O, Q, R, scales in crown view; E, N, P, scales in lateral view; A, Pi 7510; B, Pi 6808; C, Pi 7508; D, Pi 7509; E, Pi 7507; F, Pi 6806; G, Pi 7514; H, Pi 7506; I, Pi 6816; J, Pi 6809; K, Pi 7511; L, Pi 6810; M, N, Pi 7512; O, P, holotype Pi 7515; Q, Pi 6813; R, Pi 6817; S, Pi 7513; T, Pi 6811; sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: 0.1 mm.
FIG. 14 in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 14. — Set of scales of Thelodus calvus n. sp.; A, B, LIG 35-180; C, D, LIG 35-181; E, LIG 35-182; F, LIG 35-183; G, LIG 35-184; H, LIG 35-185; I, LIG 35-186; J, LIG 35-187; K, L, LIG 35-188; M, LIG 35-191; N, LIG 35-192; O, P, LIG 35-189; Q, LIG 35-190; R, LIG 35-193; sample 32a-208, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Scale bar: 0.3 mm.
FIG. 9. — A-D, Paralogania klubovi n in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 9. — A-D, Paralogania klubovi n. sp.; A, B, head scales in base view (see Fig. 8A, B); C, trunk scale of type A in base view (see Fig. 8D); D, trunk scale of type B in base view (see Fig. 8F); A, LIG 35-085; B, LIG 35-086; C, LIG 35-087; D, LIG 35-088; E, Loganelliidae gen. et sp. indet.; scale in base view (see Fig. 8J, K), LIG 35-089; sample 2zh, Pioneer Island, middle(?) Llandovery, Lower Silurian. Scale bar: 0.1 mm.
FIG. 3 in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 3. — Set of scales of Stroinolepis maenniki n. gen., n. sp., the same scales are presented in Fig. 2; A-M, Stroinolepis maenniki n. gen., n. sp.; A, head scale; B-E, transitional scales; F-M, undivided trunk and fin scales; A-G, I, L, M, scales in crown view; H, lateral oblique view; J, K, lateral view; A, Pi 7476; B, Pi 7479; C, Pi 7477; D, Pi 7484; E, Pi 7480; F, Pi 7472; G, Pi 7478; H, I, Pi 7481; J, Pi 7474, scale is lost; K, Pi 7470; L, Pi 7469; M, Pi 7473; A, C, F, G, J-M, sample MF 49-7, Strojnaya River, Ozernaya Formation, Middle-Upper Ordovician; B, E, H, I, sample MF 49-9; D, sample MF 49-11, Strojnaya River, October Revolution Island, Strojnaya Formation, Upper Ordovician. Scale bar: 0.5 mm.
FIG. 2. — A-M, Stroinolepis maenniki n. gen., n in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 2. — A-M, Stroinolepis maenniki n. gen., n. sp.; A, head scale; B-E, transitional scales; F-M, undivided trunk and fin scales; A-G, I, L, M, scales in crown view; H, lateral oblique view; J, K, lateral view; A, Pi 7476; B, Pi 7479; C, Pi 7477; D, Pi 7484; E, Pi 7480; F, Pi 7472; G, Pi 7478; H, I, Pi 7481; J, Pi 7474, scale is lost; K, Pi 7470; L, Pi 7469; M, Pi 7473; A, C, F, G, J-M, sample MF 49-7, Strojnaya River, Ozernaya Formation, Middle-Upper Ordovician; B, E, H, I, sample MF 49-9; D, sample MF 49-11, Strojnaya River, October Revolution Island, Strojnaya Formation, Upper Ordovician; N-S, Loganellia grossi Fredholm, 1990; N, head scale; O, transitional scale; P-S, trunk scales; N, O, Q-S, scales in crown view; P, a scale in base view; N, Pi 7530; O, Pi 6839; P, Pi 7533; Q, Pi 7532; R, Pi 6837; S, Pi 7535; sample 2-68, Matusevich River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Scale bars: 0.1 mm.
FIG. 6 in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 6. — Microstructure of the scales; A, B, Loganellia matura n. sp.; C-F, Shielia multispinata n. sp.; G, Paralogania consimilis n. sp.; A, Pi 7624; B, Pi 7625; C, Pi 7626; D, Pi 7627; E, Pi 7628; F, Pi 7629; G, Pi 7630; A, B, sample MF 3-8, Srednii Island, Golomyannyj Formation, middle part of Llandovery, Lower Silurian; C-F, sample MF 157-7, Spokojnaya River, upper part of Samojlovich Formation, Wenlock, Lower Silurian; G, sample MF 46-23, Ushakov River, October Revolution Island, upper part of Samojlovich Formation, Wenlock, Lower Silurian. Abbreviations: asp, aspidine "pegs" in the base; b, base; bc, canal in the base; cr, crown; dc, dentine canal; dt, dentine tubules; fpc, fan-shaped pulp canals; lac, lacunae of dentine canals; mpc, main pulp canal; n, neck; pc, pulp canal; pco, pulp canal opening; sp, spine. Scale bars: 0.1 mm.
FIG. 10. — Paralogania klubovi n in Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia)
FIG. 10. — Paralogania klubovi n. sp., microstructure of the scales; A, head scale, LIG 35-086; B, trunk scale, LIG 35-087; the scales were drawn in anise oil; sample 2zh, Pioneer Island, middle(?) Llandovery, Lower Silurian. Abbreviations: b, base; cr, crown; dt, dentine tubules; odt, openings of dentine tubules; mpc, main pulp canal. Scale bar: 0.1 mm.
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