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zenodo32/100

FIGURES 18 a–o in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 18 a–o. Male habitus in dorsal view of: H. lazica (Turkey, Giresun, Kümbet) (a); H. hosseinieorum (Iran, Fars, N Dalin) (b); H. berthelemyana (From Jäch et al. 2006: Lebanon) (c). Aedeagi in right side view of: H. lazica (Turkey, Giresun, Kümbet) (d); H. hosseinieorum (Iran, Fars, N Dalin) (e); H. berthelemyana (From Jäch et al. 2006: Lebanon, Abou Ali) (f); H. orthosia (From Jäch et al. 2006: Lebanon, El Bared, Sir el Danniyeh) (g); H. fontiscarsavii (Turkey, Konya, 6 km E Kösere) (h). Female elytral terminations of: H. lazica (Turkey, Giresun, Kümbet) (l); H. hosseinieorum (Iran, Fars, N Dalin) (m); H. berthelemyana (Lebanon, Abou Ali) (n); H. orthosia (Lebanon, El Bared, Sir el Danniyeh) (o); H. fontiscarsavii (Turkey, Konya, 6 km E Kösere) (p). Scale bar = 1 mm (Figs. 18a–c); = 0.25 mm (Figs. 18d–h). Female elytral terminations not scaled.

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zenodo32/100

FIGURES 20 a–l in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 20 a–l. Male habitus in dorsal view of: H. planata (From Jäch &amp; Díaz 2006: Azerbaijan, Gäncä) (a). Aedeagi in right side view of: H. planata (From Jäch &amp; Díaz 2006: Azerbaijan, Gäncä) (b); H. solodovnikovi (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, NE Dagomys) (c); H. krasnodarensis (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, 13 km ENE Sotchi) (d); H. prokini (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, Novorossiysk) (e). Female elytral terminations of: H. planata (From Jäch &amp; Díaz 2006: Azerbaijan, Gäncä) (f); H. solodovnikovi (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, NE Dagomys) (g); H. krasnodarensis (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, 13 km ENE Sotchi) (h); H. prokini (From Jäch &amp; Díaz 2006: Russia, Krasnodarskiy Kray, Novorossiysk) (l). Scale bar = 1 mm (Figs. 20a); = 0.25 mm (Figs. 20b–e). Female elytral terminations not scaled.

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zenodo32/100

FIGURES 12 a–m in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 12 a–m. Male habitus in dorsal view of: H. iberica (Spain, Salamanca, Candelario) (a); H. lusitana (Portugal) (b). Aedeagi in right side view of: H. iberica (Spain, Salamanca, Candelario) (c); H. altamirensis (From Díaz Pazos &amp; Garrido Gonzalez 1993: Spain, Extremadura, Cáceres) (d); H. madronensis (From Castro et al. 2000: Spain, Castilla, Fuencaliente) (e); H. lusitana (Portugal) (f). Female elytral terminations of: H. iberica (Spain, Salamanca, Candelario) (g); H. altamirensis (Spain, Extremadura, Cáceres) (h); H. madronensis (Spain, Castilla, Fuencaliente) (l); H. lusitana (Portugal) (m). Scale bar = 1 mm (Figs. 12a–b); = 0.25 mm (Figs. 12c–f). Female elytral terminations not scaled.

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FIGURES 16 a–q in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 16 a–q. Male habitus in dorsal view of: H. caucasica (Turkey, Rize, Ikizdere) (a); H. septemlacuum (Turkey, Bolu, Yedigöller N. P.) (b). Aedeagi in right side view of: H. caucasica (Turkey, Rize, Ikizdere) (c); H. septemlacuum (Turkey, Bolu, Yedigöller N. P.) (d); H. sinope (Turkey, Sinop, Dranaz pass) (e); H. plastica (Turkey, Bursa, Keschish dag) (f); H. khnzoriani (Turkey, Tatvan, Mutki) (g); H. nilguenae (Turkey, Adana, Gobelli) (h). Female elytral terminations of: H. caucasica (Turkey, Rize, Ikizdere) (l); H. septemlacuum (Turkey, Bolu, Yedigöller N. P.) (m); H. sinope (Turkey, Sinop, Dranaz pass) (n); H. plastica (Turkey, Bursa, Keschish dag) (o); H. khnzoriani (Turkey, Tatvan, Mutki) (p); H. nilguenae (Turkey, Adana, Saimbeyli) (q). Scale bar = 1 mm (Figs. 16a–b); = 0.25 mm (Figs. 16c–h). Female elytral terminations not scaled.

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FIGURE 49 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 49. Geographic distribution of: H. dentipes; H. heterogyna; H. plumipes; H. discreta; H. producta.

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FIGURE 33 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 33. Geographic distribution of: H. excisa; H. akbesiana; H. integra; H. cata; H. phallica; H. christinae; H. exasperata.

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zenodo32/100

FIGURE 19 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 19. Geographic distribution of: H. lazica; H. hosseinieorum; H. berthelemyana; H. plastica; H. orthosia; H. fontiscarsavii.

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zenodo32/100

FIGURES 48 a–s in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 48 a–s. Male habitus in dorsal view of: H. dentipes (Italy, Trentino, Bieno) (a); H. heterogyna (Italy, Toscana, Pescina) (b); H. plumipes (Italy, Toscana, Pracchia) (c); H. discreta (Italy, Liguria, Amborzasco) (d). Aedeagi in right side view of: H. dentipes (Italy, Trentino, Bieno) (e); H. heterogyna (From Audisio &amp; De Biase 1995: Italy, Lombardia, Stazzona) (f); H. plumipes (Italy, Toscana, Piancaldoli) (g); H. discreta (Italy, Emilia, Groppallo) (h); H. producta (France, Languedoc, Aude) (l). Female elytral terminations of: H. dentipes (Italy, Lombardia, Dazio) (m); H. heterogyna (Italy, Toscana, Orsigna; Italy, Lucania, Tito; Italy, Liguria, Fiorino) (n,o,p); H. plumipes (Italy, Toscana, Pratomagno) (q); H. discreta (Italy, Liguria, Amborzasco) (r); H. producta (France) (s). Scale bar = 1 mm (Figs. 48a–d); = 0.25 mm (Figs. 48e–l). Female elytral terminations not scaled.

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FIGURES 50 a–b in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 50 a–b. Male habitus in dorsal view of: H. sappho (Holotype: Greece, Lefkas) (a). Aedeagus in right side view of: H. sappho (Holotype: Greece, Lefkas—aedeagus probably damaged) (b). Scale bar = 1 mm (Fig. 50a); = 0.25 mm (Fig. 50b).

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zenodo32/100

FIGURES 40 a–m in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 40 a–m. Male habitus in dorsal view of: H. devillei (Italy, Latium, Tolfa) (a); H. lapidicola (Italy, Piemonte, Ala di Stura) (b). Aedeagi in right side view of: H. devillei (Italy, Latium, S. Vittorino) (c); H. devincta (Italy, Veneto, Collalto) (d); H. lapidicola (From Audisio &amp; De Biase 1995: Italy, Liguria, Colle del Melogno) (e); H. sanfilippoi (From Audisio &amp; De Biase 1995: Italy, Liguria, Fiorino) (f). Female elytral terminations of: H. devillei (Italy, Toscana, Vivo d'Orcia) (g); H. devincta (Italy, Friuli, Attimis) (h); H. lapidicola (Italy, Trentino, Bieno) (l); H. sanfilippoi (Italy, Liguria, Fiorino) (m). Scale bar = 1 mm (Figs. 40a–b); = 0.25 mm (Figs. 40c–f). Female elytral terminations not scaled.

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FIGURES 46 a–p in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 46 a–p. Male habitus in dorsal view of: H. bensae (Italy, Liguria, Nava) (a); H. polita (France, Provence, Vence) (b); H. bicuspidata (France, Rhône, St. Romain en Giers) (c). Aedeagi in right side view of: H. bensae (France, Provence, La Bolléne) (d); H. polita (Austria, Tyrol, Steinach) (e); H. bicuspidata (France, Rhône, St. Romain en Giers) (f); H. bitruncata (From Fresneda et al. 1994: France, Languedoc, La Preste) (g); H. catalonica (Spain, Catalonia, El Burgassos) (h). Female elytral terminations of: H. bensae (Italy, Liguria, Nervia river) (l); H. polita (Austria, Tyrol, Steinach) (m); H. bicuspidata (France, Rhône, St. Romain en Giers) (n); H. bitruncata (Spain, Catalonia, Gerona) (o); H. catalonica (Spain, Catalonia, El Burgassos) (p). Scale bar = 1 mm (Figs. 46a–c); = 0.25 mm (Figs. 46d–h). Female elytral terminations not scaled.

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FIGURE 45 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 45. Geographic distribution of: H. tatii; H. manfredjaechi; H. gaditana; H. monstruosipes; H. zezerensis.

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FIGURES 32 a–r in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 32 a–r. Male habitus in dorsal view of: H. akbesiana (Turkey, Adana, Yarpuz) (a). Aedeagi in right side view of: H. excisa (Austria, Donnerskirchen) (b); H. akbesiana (From Audisio et al. 1993: Turkey, Adana, Yarpuz) (c); H. integra (From Audisio et al. 1996: Turkey, Artvin, Borçka) (d); H. cata (From Audisio et al. 1996: Turkey, Çanakkale) (e); H. phallica (From Audisio et al. 1996: Greece, Etolia, Pentakorfos) (f); H. christinae (From Audisio et al. 1996: Greece, Peloponnesus, Kato Vlasia) (g); H. exasperata (Spain, Castilla, Alcaraz) (h). Female elytral terminations of: H. excisa (Rumania, Ilba) (l); H. integra (Turkey, Istanbul, NE Teke) (m); H. akbesiana (Complete Female habitus: Turkey, Adana, Çatak) (n); H. cata (Turkey, Mugla, Kale) (o); H. phallica (Greece, Evritania, Karpenissi) (p); H. christinae (Greece, Peloponnesus, Kato Vlasia) (q); H. exasperata (Spain, Leon, Pedrosa del Rey) (r). Scale bar = 1 mm (Figs. 32a,n); = 0.25 mm (Figs. 32b–h). Female elytral terminations not scaled.

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FIGURES 30 a–h in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 30 a–h. Male habitus in dorsal view of: H. samnitica (Italy, Abruzzo, Orfento river) (a); H. pelops (Greece, Peloponnesus, Vasiliki—Gorani) (b). Aedeagi in right side view of: H. samnitica (Italy, Abruzzo, Pietracamela) (c); H. pangaei (From Audisio et al. 1996: Greece, Kavala, Pangaeon Mt.) (d); H. pelops (Greece, Peloponnesus, Vasiliki—Gorani) (e). Female elytral terminations of: H. samnitica (Italy, Abruzzo, Pietracamela) (f); H. pangaei (Greece, Kavala, Pangaeon Mt.) (g). Female habitus in dorsal view of H. pelops (Greece, Peloponnesus, Vasiliki—Gorani) (h). Scale bar = 1 mm (Figs. 30a,b,h); = 0.25 mm (Figs. 30c–e). Female elytral terminations not scaled.

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FIGURES 42 a–g in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 42 a–g. Male habitus in dorsal view of: H. bosnica (Bosnia, Sarajevo) (a). Aedeagi in right side view of: H. bosnica (From Audisio et al. 1996: Bosnia, Sarajevo) (b); H. hungarica (Ukraine, Gyertyánliget) (c); H. leonhardi (From Audisio et al. 1996: Greece, Rodopi, Dipotama) (d). Female elytral terminations of: H. bosnica (Bosnia, Budmolici) (e); H. hungarica (Ukraine, Gyertyánliget) (f); H. leonhardi (Greece, Rodopi, Dipotama) (g). Scale bar = 1 mm (Fig. 42a); = 0.25 mm (Figs. 42b–d). Female elytral terminations not scaled.

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FIGURE 4 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 4. Fastflowing perennial stream with surrounding vegetation, typical habitat of a few "Haenydra" species.

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FIGURES 22 a–l in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 22 a–l. Male habitus in dorsal view of: H. vedrasi (Greece, Evritania, Karpenissi) (a); H. jaechiana (Peloponnesus, Tripoli, Daras) (b); H. epeirosi (Greece, Fthiotida, Gardiki) (c). Aedeagi in right side view of: H. vedrasi (From Audisio et al. 1996: Greece, Evritania, Karpenissi) (d); H. jaechiana (From Audisio et al. 1996: Peloponnesus, Tripoli, Daras) (e); H. epeirosi (From Audisio et al. 1996: Greece, Fthiotida, Gardiki) (f). Female elytral terminations of: H. vedrasi (Greece, Evritania, Anatholios-Fragista) (g); H. jaechiana (Peloponnesus, Tripoli, Castorio) (h); H. epeirosi (Greece, Etolia, Pentakorfos) (l). Scale bar = 1 mm (Figs. 22a–c); = 0.25 mm (Figs. 22d–f). Female elytral terminations not scaled.

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FIGURES 26 a–u in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 26 a–u. Male habitus in dorsal view of: H. larissae (Italy, Piemonte, Oropa) (a); H. hispanica (Spain, Salamanca, Candelario) (b). Aedeagi in right side view of: H. emarginata (Spain, Leon, Puerto S. Isidro) (c); H. larissae (Italy, Trentino, Grigno; Italy, Piemonte, Oropa) (d–e); H. diazi (Spain, Guipúzcoa, Oiartzun) (f); H. fosterorum (Spain, Burgos, Fresneda de la Sierra) (g); H. hispanica (Spain, Salamanca, Candelario) (h); H. saga (Bosnia, Visegrad) (l); H. alpicola (Slovenia, Valdekom) (m). Female elytral terminations of: H. emarginata (Spain, Leon, Puerto S. Isidro) (n); H. larissae (Italy, Trentino, Stava; Italy, Piemonte, Sagliano) (o–p); H. diazi (Spain, Guipúzcoa, Oiartzun) (q); H. fosterorum (Spain, Burgos, Fresneda de la Sierra) (r); H. hispanica (Spain, Leon, Valdo Prado) (s); H. saga (Slovakia, Muran) (t); H. alpicola (Austria, Scheibbs) (u). Scale bar = 1 mm (Figs. 26a–b); = 0.25 mm (Figs. 26d–m). Female elytral terminations not scaled.

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FIGURES 10 a–n in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURES 10 a–n. Male habitus in dorsal view of: H. schuleri (Slovenia, Zidanl) (a); H. subintegra (Greece, Evritania, Karpenissi) (b); H. carniolica (Italy, Veneto, Falzè di Piave) (c). Aedeagi in right side view of: H. schuleri (Slovenia, Zidanl) (d); H. subintegra (Greece, Evritania, Karpenissi) (e); H. aroensis (Peloponnesus, Kato Vlasia) (f); H. carniolica (Italy, Veneto, Falzè di Piave) (g). Female elytral terminations of: H. schuleri (Italy, Friuli, Cosizza river) (h); H. subintegra (Greece, Evritania, Karpenissi) (l); H. aroensis (Peloponnesus, Kato Vlasia) (m); H. carniolica (Italy, Veneto, Falzè di Piave) (n). Scale bar = 1 mm (Figs. 10a–c); = 0.25 mm (Figs. 10d–g). Female elytral terminations not scaled.

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FIGURE 9 in <strong>A revision of <em>Hydraena</em> species of the "<em>Haenydra</em>" lineage (Coleoptera, Hydraenidae)</strong>

FIGURE 9. Used male genitalia terminology. Aedeagus in right side view of H. epeirosi (Greece, Fthiotida, Gardiki).

opennotspecifiedJan 2013View details →

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