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zenodo32/100

FIGURE 3 in Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan

FIGURE 3. Adult of Rhyacophila bifue sp. nov. 3A–3D, male: 3A, genitalia, left lateral, omitting left inferior appendage; 3B, same, dorsal; 3C, right inferior appendage, dorsal; 3D, same, ventral. 3E–3I, female: 3E, abdominal segments VIII–XI, left lateral; 3F, segment VIII, dorsal; 3G, same, ventral; 3H, vaginal apparatus, left lateral; 3I, same, ventral. Abbreviations: VIII–XI = abdominal segments VIII–XI; a inf app = apical segment of inferior appendage (paired); ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); cu ri = curved ridge (paired); do me pr = dorsomesal process (paired); pp = posterior process; ps = processus spermathecae; ru po IX = rugged posterior area of segment IX; te st = tergal strap (paired).

opennotspecifiedNov 2024View details →
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FIGURE 2 in Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan

FIGURE 2. Adult of Rhyacophila eniwa sp. nov. 2A–2D, male: 2A, genitalia, left lateral, omitting left inferior appendage; 2B, same, caudal, omitting inferior appendages; 2C, dorsal branch of segment X, dorsal; 2D, right inferior appendage, ventral. 2E–2I, female: 2E, abdominal segments VIII–XI, left lateral; 2F, segment VIII, dorsal; 2G, same, ventral; 2H, vaginal apparatus, left lateral; 2I, same, ventral. Abbreviations: VIII–XI = abdominal segments VIII–XI. a inf app = apical segment of inferior appendage (paired); ae = aedeagus; ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); do br X = dorsal branch of segment X; do pr = dorsal process of phallobase; par = paramere; pp = posterior process; ps = processus spermathecae; ve br X = ventral branch of segment X; ve lo = ventral lobe of aedeagus; ve pr = ventral process of phallobase (paired).

opennotspecifiedNov 2024View details →
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FIGURE 1 in Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan

FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B–1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori & Ohkawa; 1E, R. shimamaki sp. nov.

opennotspecifiedNov 2024View details →
zenodo32/100

FIGURES 9–10 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURES 9–10. Wings of Rhyacophila fasciata Hagen 1859 and Rhyacophila anatolica Ekingen & Valladolid sp. nov. 9a, forewing of R. fasciata, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 9b, forewing of R. fasciata male from Slovakia; 9c, hind wing of R. fasciata male from Slovakia. 10a, forewing of R. anatolica male. 10b, hind wing of R. anatolica male. St = stigma. Major veins: C = costa, Sc = subcosta, R 1 –R = 5 radius, M 1 –M 4 = media, Cu1–Cu2 = cubitus, A1–A3 = anal; m-cu = crossvein between medial and cubital veins, cu-a = crossvein between second cubital and first anal veins (from Holzenthal et al. 2007). Scale bars: 2 mm. 9a: specimen preserved dry, 9b–c, 10a–b: specimens preserved in ethanol.

opennotspecifiedNov 2024View details →
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FIGURES 15–18 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen & Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm.

opennotspecifiedNov 2024View details →
zenodo32/100

FIGURE 8 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURE 8. Rhyacophila anatolica Ekingen & Valladolid sp. nov., paired abdominal hook plates of male pupa. 8l and 8r, left and right hook plates, respectively. A = anterior hook plates, dorsal, P = posterior hook plates, dorsal; III–VII = abdominal terga III through VII, dorsal. Scale bars: 50 μm.

opennotspecifiedNov 2024View details →
zenodo32/100

FIGURE 1 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURE 1. Setae (S) and sensory pits (P) of the head and thorax of Rhyacophila larvae. 1a, head, dorsal; 1b, prothorax, right sclerite, dorsal (from Williams & Wiggins 1981).

opennotspecifiedNov 2024View details →
zenodo32/100

FIGURES 11–12 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen & Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm.

opennotspecifiedNov 2024View details →
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FIGURES 2–7 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen & Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm.

opennotspecifiedNov 2024View details →
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FIGURE 20 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURE 20. Spatial distribution of Rhyacophila anatolica Ekingen & Valladolid sp. nov. in Turkey. Star: specimens with DNA information. B. Bulgaria, C: Cyprus, Ge: Georgia, Gr: Greece, I: Iraq, R: Russia, S: Syria.

opennotspecifiedNov 2024View details →
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FIGURES 13–14 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen & Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm.

opennotspecifiedNov 2024View details →
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FIGURE 19 in The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen & Valladolid, sp. nov., based on morphological, genetic, and ecological evidence

FIGURE 19. Phylogenetic relationships among the species of the "Rhyacophila fasciata Species Complex" included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid & Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid & Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid & Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić & Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky & Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch.

opennotspecifiedNov 2024View details →
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FIGURES 1–3. Rhyacophila spp., male. 1A–1F in Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan

FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX.

opennotspecifiedNov 2021View details →
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FIGURE 9 in Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan

FIGURE 9. Distributions of 4 Rhyacophila species in East Asia. Distribution areas of R. confissa and R. vicina are cited from Ivanov (2011), Park & Kong (2020), and Yang et al. (2016).

opennotspecifiedNov 2021View details →
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FIGURE 19 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURE 19. Spatial distribution of Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.), Rhyacophila kykladica Malicky & Sipahiler 1993 and Rhyacophila fasciata Hagen 1859 in Greece, Republic of North Macedonia, Kosovo and Serbia. Circle: localities of R. macedonica. Triangles: localities of R. fasciata. Diamond: localities of R. kykladica. White, orange: specimens from Greece. Green: specimens from Kosovo. Purple: specimens from Republic of North Macedonia. Brown: specimens from Serbia.

opennotspecifiedApr 2022View details →
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FIGURES 10, 11 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURES 10, 11. Male and female genitalia of Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.), respectively. 10, male: 10A, 2nd segment of left inferior appendage, left lateral; 10BV, 10BL, parameres: 10BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 10BL, left paramere, left lateral. 10CL, 10CD, aedeagus (phallicata) and lateroventral lobes: 10CL, aedeagus and ventral lobe, left lateral; 10CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa); 10DV, segment X, ventral, ab= apical band, as= anal sclerites, va= nonsclerotized ventral area. 11, female segments VIII–XI: 11L, left lateral; 11D, dorsal; 11V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 µm.

opennotspecifiedApr 2022View details →
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FIGURES 8, 9 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURES 8, 9. Wings of Rhyacophila fasciata Hagen 1859 and Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.). 8a, forewing of R. fasciata, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 8b, forewing of R. fasciata male from Kosovo; 8c, hind wing of R. fasciata male from Kosovo. 9a, 9b, forewings of R. macedonica males from Kosovo; 9c, hind wing of R. macedonica male from Kosovo. Major veins: C = costa, Sc = subcosta, R1–R 5 = radius, M1–M4 = media, Cu1–Cu2 = cubitus, A1–A3 = anal, m-cu = crossvein between medial and cubital veins (from Holzenthal et al. 2007). Scale bars: 2 mm.

opennotspecifiedApr 2022View details →
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FIGURES 14–17 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURES 14–17. Males and females of Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.) and Rhyacophila fasciata Hagen 1859 from Kosovo. 14, male of R. macedonica: 14a, 2nd segment of left inferior appendage, left lateral; 14b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 14c, parameres and ventral lobe of aedeagus (phallicata), ventral; 14d, left paramere, left lateral. 15, male of R. fasciata: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, female segments VIII–XI of R. macedonica: 16L, left lateral; 16D, dorsal; 16V, ventral. 17, female segments VIII–XI of R. fasciata: 17L, left lateral; 17D, dorsal; 17V, ventral. Scale bars: Figures 14a–c, 15a–c = 0.5 mm, Figures 14d, 15d = 200 µm, Figures 16, 17 = 1 mm.

opennotspecifiedApr 2022View details →
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FIGURES 12, 13 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURES 12, 13. Larvae of Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.) and Rhyacophila fasciata Hagen 1859. 12, R. macedonica: 12a, head, dorsal; 12b, prothorax, dorsal; 12c, abdominal tergite IX, dorsal; 12d, left anal proleg, left lateral. 13, R. fasciata: 13a, head, dorsal. 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. Scale bars: 1 mm.

opennotspecifiedApr 2022View details →
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FIGURE 7 in The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae)

FIGURE 7. Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.), paired abdominal hook plates of male pupa. 7l and 7r, left and right hook plates, respectively. A = anterior hook plates, dorsal, P = posterior hook plates, dorsal; III–VII = abdominal terga III through VII, dorsal. Scale bars: 50 µm.

opennotspecifiedApr 2022View details →

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