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FIGURE 8. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 8. Hyalella cuyana n. sp. (photograph), female paratype (FML-CRUST 01284), embryos in brood pouch.

FIGURE 9. A in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 9. A Photographs and Google Earth satellite image of sampling sites (s) at the Calingasta River. Type locality of Hyalella cuyana n. sp. on the year 2014 B Photographs of Calingasta River in the vicinity of the sampling site taken in 2015.

FIGURE 5. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 5. Hyalella cuyana n. sp., male holotype (FML-CRUST 01282) and female paratype (FML-CRUST 01284). A male gnathopod 1, arrows indicate enlargement of the palmar angle B male gnathopod 2 C female gnathopod 2, arrows indicate curled setae of oostegites. Scale bars: 0.1mm.

FIGURE 2 in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 2. Habitus (photograph) of Hyalella cuyana n. sp. A female paratype (FML-CRUST 01284) B male holotype (FMLCRUST 01282).

FIGURE 7. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 7. Hyalella cuyana n. sp., male holotype (FML-CRUST 01282). A pleopod 3 B uropod 1 C uropod 2 D uropod 3 E telson of male paratype F telson lateral view. Scale bars: 0.1mm.

FIGURE 6. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 6. Hyalella cuyana n. sp., male holotype (FML-CRUST 01282). A pereiopod 3 B pereiopod 4 C pereiopod 5 D pereiopod 6 E pereiopod 7.

FIGURE 1 in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 1. Map showing distribution of type locality of Hyalella cuyana n. sp. and records of Cuyojanira riojana, Cuyojanira ischicuca, Cuyojanira sp. (Isopoda), Parastygocaris andina (Anaspidacea), Hyalella rionegrina (Amphipoda) in Argentina within the biogeographic scheme of Arana et al. (2017).

FIGURE 11 in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 11. Habitus (photograph) Parastygocaris andina (FML-CRUST 01292) (Anaspidacea, Stygocarididae).

FIGURE 4. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 4. Hyalella cuyana n. sp., male holotype (FML-CRUST 01282). A left maxilliped B labrum C lower lip D left mandible E right mandible F internal lateral view of maxilla 1 G maxilla 1 H maxilla 2. Scale bars: 0.1 mm.

FIGURE 3. Hyalella cuyana n in South America stygobiont crustaceans: a new species of Hyalellidae (Amphipoda) and new reports of Stygocarididae (Anaspidacea) and Protojaniridae (Isopoda) from Calingasta Valley, Pre-Andean region, San Juan, Argentina

FIGURE 3. Hyalella cuyana n. sp., male holotype (FML-CRUST 01282). A paratype male (FML-CRUST 01283), habitus lateral view B Epimeral plates C antenna 1 D antenna 2

Fig. 9 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 9 (next page). A. Plagigeyeria erossi sp. nov., Mostar district, Komadinovo Vrelo at right bank of Neretva below Donja Jablanica, holotype (HNHM-MOLL- 104166). B–C. P. erossi sp. nov., type locality, paratypes (JG F0518, JG F1122). D. P. lewarnei sp. nov., Trebinje district, Gorica, cave Vruljak, holotype (HNHM-MOLL-104176). E. P. listicaensis sp. nov., Široki Brijeg, Bilo Vrelo (Lištica 2), holotype (HNHM-MOLL-104168). F. P. listicaensis sp. nov., type locality, paratype (JG F1250). G–H. P. reischuetzorum sp. nov., Dabarsko Polje, Vrelo Vrijeka. G. Paratype (HNHM-MOLL-104165). H. Paratype (JG F1173). I. P. reischuetzorum sp. nov., type locality, holotype (NHMW 111678). J. P. reischuetzorum sp. nov., type locality, paratype (JG F1158). K–L. Travunijana gloeri sp. nov., Dabarsko Polje, Vrelo Vrijeka. K. Holotype (HNHM-MOLL-104418). L. Paratype (JG F1340).

Fig. 7 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 7 (next page). A. Plagigeyeria mostarensis, Kuščer 1933, holotype, from Kuščer 1933, assigned to Izvor Buna in Blagaj but rather represents the form from near spring Bunica in Hodbina. B. P. mostarensis, hypotype of Schütt, 1970, from spring Bunica (SMF 221236). C. P. mostarensis, likely topotype from spring Bunica (JG F1146). D–E. P. mostarensis var., from spring of Buna assigned as type locality (JG F0536). Fa–Fb. Paladilhiopsis (Costellina) turrita (Kuščer, 1933) paratype and holotype from Kuščer 1933, spring Izvor in valley of river Jadro near Split. G. P. pseudocostellina sp. nov., left bank side spring of river Buna 400 m from main spring (holotype HNHM-MOLL-104164). H. P. pseudocostellina sp. nov., left bank side spring of river Buna 200 m from main spring (paratype JG F1140). I. Travunijana ovalis (Kuščer, 1933), holotype from Kuščer, assigned to Izvor Buna in Blagaj but not found in assigned locality, only in spring Bunica in Hodbina. J–K. T. ovalis (Kuščer, 1933), spring Bunica near Hodbina (JG F1145). L. T. robusta (Schütt, 1959), Trebinje district, Tučevac spring at right bank of Trebišnjica River (JG F1163). M. T. robusta, Trebinje district, Gorica, cave Vruljak 1 (JG F0831). N. T. robusta asculpta (Schütt, 1972), spring Ombla (Dubrovačka Rijeka) at Komolac near Dubrovnik (JG F1080).

Fig. 5 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 5 (next page). A. Plagigeyeria plagiostoma (A. J. Wagner, 1914), from Vrelo Bosne in Ilidža, (topotype JG F1116). B. Travunijana vruljakensis Grego & Glöer, 2019, Gorica near Trebinje, spring Studenac (holotype HNHM-MOLL-104416). C. P. inflata (A. J. Wagner, 1928), from Vrelo Bosne in Ilidža (topotype JG F1117). D. P. ljutaensis sp. nov., Konjic district, spring of River Ljuta (holotype HNHM-MOLL-104180). E. P. konjicensis sp. nov., small spring at left bank of Ljuta River, 600 m downstream of the main spring (holotype HNHM-MOLL-104174); F–G. P. olsavskyi sp. nov., Studenci, spring Kajtazovina F. Holotype (HNHM-MOLL-104178). G. Paratype (JG F1192). H–I. P. olsavskyi sp. nov., Ljubuški district, Donji Proložac, spring Mali Prokop (JG F1209).

Fig. 1 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 1 (next page). Studied localities of Plagigeyeria and Travunijana in Hercegovina and phreatic hydrology of the related karstic aquifers with flow directions of the groundwater. Hydrological situation after Stevanović et al. 2014. 1–14. Newly studied localities (red dot ●). 1. Konjic, spring Ljuta (LT of Plagigeyeria ljutaensis sp. nov.). 2. Konjic spring at left side of Ljuta stream (LT of P. konjicensis sp. nov.). 3. Donja Jablanica, Komadinovo Vrelo (LT of P. erossi sp. nov.). 4. Široki Brijeg, spring Lištica, Bilo Vrilo (LT of P. listicaensis sp. nov.). 5. Studencí, spring Kajtazovina (LT of P. olsavskyi sp. nov.). 6. Studencí, Vrilo Guljevina (LT of P. jakabi sp. nov.). 7. Donji Proboj near Ljubuški, spring Mali Prokop (P. olsavskyi sp. nov.). 8. Vitina, spring Vrioštica (LT of P. vriosticaensis sp. nov. and P. ozimeci sp. nov.). 9. Lukende, Jakšenica spring (LT of P. angyaldorkae sp. nov.). 10. Studenci, spring Vakuf (P. cf. jakabi). 11. Dabarsko Polje, spring Vrijeka (LT of P. reischuetzorum sp. nov. and Travunijana gloeri sp. nov.). 12. Trebinje, spring Tučevac (T. robusta, Schütt, 1959 and P. cf. lewarnei sp. nov.). 13. Trebinje, Gorica, spring Vruljak (T. robusta and LT of P. lewarnei sp. nov.). 14. Vučija, Sušićka Jama Estavelle (P. lewarnei sp. nov.). 15–32. Historical localities (blue dot ●): 15. Blagaj, spring of Buna (P. mostarensis var. Kuščer, 1933 and LT of P. pseudocostellina sp. nov.). 16. Hodbina, spring of Bunica (P. mostarensis Kuščer, 1933 and Travunijana ovalis (Kuščer, 1933)). 17. Bileća, spring Trebišnjica (T. robusta (Schütt, 1959) and T. tribunicae (Schütt, 1953)). 18. Čepelica, spring Čepo (T. robusta (Schütt, 1959)). 19. Bileća, spring Miruše (T. robusta (Schütt, 1959) and T. tribunicae (Schütt, 1953), P. zetatridyma Schütt, 1959). 20. Bileća, spring Dobriševo (T. robusta, (Schütt, 1959)). 21. Fatničko Polje, Obod Cave (T. robusta (Schütt, 1959), P. reischuetzorum sp. nov., T. gloeri sp. nov.). 22. Gacko, Cave Stepen (T. robusta). 23. Mlini, spring south of Plat (T. robusta asculpta Schütt, 1972). 24. Komolac, spring Ombla (T. robusta asculpta Schütt, 1972 and T. angelovi Schütt, 1972,). 25. Mali Zaton, spring Mlin (T. robusta asculpta (Schütt, 1972), T. angelovi (Schütt, 197)). 26. Svitavsko Blato, spring Sjekoše (T. edlaueri (Schütt, 1961), T. nitida (Schütt, 1963)). 27. Svitavsko Blato, spring Sopot Mlin (T. edlaueri (Schütt, 1961), T. nitida (Schütt 1963)). 28. Svitavsko Blato, spring Bajovci (T. edlaueri (Schütt, 1961)). 29. Metković, spring Glušči (T. edlaueri and T. nitida (Schütt, 1963)). 30. Ploče, spring Ocuša at bank of Bačinsko Jezero Lake (T. klemmi (Schütt, 1961)). 31. Vrgorac, spring Stenjevac (T. klemmi (Schütt, 1961)). 32. Ilidža, Vrelo Bosne (P. plagiostoma (A. J. Wagner, 1914) and P. inflata (A. J. Wagner, 1928)). 1–43. Karst basins – Poljes of the studied region (green square ■): 1. Konavosko Polje, 50 m a.s.l. 2. Zubačko Polje, 672 m a.s.l. 3. Konjsko and Carevo P., 829–875 m a.s.l. 4. Mokro Polje, (Popovo Polje) 269 m a.s.l. 5. Popovo Polje, 227 m a.s.l. 6. Jasen Polje, 556 m a.s.l. 7. Ljubomirsko Polje, 506 m a.s.l. 8. Ljubinjsko Polje, 396 m a.s.l. 9. Plansko Polje, 609 m a.s.l. 10. Fatničko Polje, 452 m a.s.l. 11. Dabarsko Polje, 472 m a.s.l. 12. Lukavačko Polje, 895 m a.s.l. 13. Slato Polje, 1012 m a.s.l. 14. Cerničko Polje, 816 m a.s.l. 15. Gatačko Polje, 936 m a.s.l. 16. Nevesinjsko Polje, 817 m a.s.l. 17. Trusinsko Polje, 866 m a.s.l. 18. Stolačko Polje, 45 m a.s.l. 19. Rotimlja Polje, 422 m a.s.l. 20. Crničko Polje, 212 m a.s.l. 21. Hutovo Blato, 2 m a.s.l. 22. Svitavsko Blato, 2 m a.s.l. 23. Gradac Polje, 88 m a.s.l. 24. Vrgorsko (Jezerac) Polje, 20 m a.s.l. 25. Rastok Polje, 72 m a.s.l. 26. Ljubuško Polje, 58 m a.s.l. 27. Studeničko Polje, 25 m a.s.l. 28. Čitluško Polje, 217 m a.s.l. 29. Mostarsko Blato, 223 m a.s.l. 30. Mokro Polje, 230 m a.s.l. 31. Rašanjsko Polje, 344 m a.s.l. 32. Kočerinsko Polje, 302 m a.s.l. 33. Imotsko & Grudsko P., 251–280 m a.s.l. 34. Posuško Polje, 578 m a.s.l. 35. Dugo (Dugorudo) Polje, 1206 m a.s.l. 36. Ravanjsko Polje, 1131 m a.s.l.; 37. Rakitno Polje, 890 m a.s.l. 38. Vučipolje, 977 m a.s.l.; 39. Roško Polje, 894 m a.s.l. 40. Duvanjsko Polje, 865 m a.s.l. 41. Džepi Polje, 830 m a.s.l. 42. Prečko Polje, 1002 m a.s.l. 43. Hansko Polje, 835 m a.s.l. 44. Donje Ziemlje, 507 m a.s.l.

Fig. 19 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 19. Localities of species of Travunijana in Hercegovina and adjacent regions and the most likely distribution range of the known species within the related aquifers (colour blotches) and sampling localities (dots with numbers). Freshwater springs are in circles and brackish or submarine springs are as marked by diamonds, arrows show main karst conduits and locality numbers are according to the legend from Fig. 1.

Fig. 14. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 14. A. Travunijana vruljakensis Grego & Glöer, 2019, Hercegovina, Vrelo Studenac, Gorica near Trebinje, SBMNH 625961 (SEM SBMNH Vanessa Delnavaz). B. T. angelovi (Schütt, 1972), Spring Ombla, Komolac near Dubrovnik, paratype (SMF 318892). C–D. T. tribunicae (Schütt, 1963), Hercegovina, Bileća, main spring of Trebijšnica. C. SMF 191518. D. SMF 318909. Scale bars = 1 mm (B–D SEM SMF Sigrid Hof).

Fig. 16. A–B in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 16. A–B. Travunijana klemmi (Schütt, 1961), Croatia, spring in Stenjevac near Vrgorac. A. SMF 318873. B. Paratype SMF 164345. C. T. edlaueri (Schütt, 1961), Hercegovina, Sopot Mlin at south bank of Svitavsko Blato near Metkovic, SMF 171671. D. T. edlaueri, Hercegovina, Sjekoše at south bank of Svitavsko Blato near Metkovic, SMF 318858. Scale bars = 1 mm (SEM SMF Sigrid Hof).

Fig. 11. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 11. A. Plagigeyeria ozimeci sp. nov., Vitina, Vrilo Vrioštica, paratype SBMNH 626440. B. Plagigeyeria lewarnei sp. nov., spring Tučevac in Trebinje, SBMNH 626406. Scale bars = 1 mm (SEM SBMNH Vanessa Delnavaz).

Fig. 8. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 8. A. Plagigeyeria reischuetzorum sp. nov., Bjeljani, Dabarsko Polje, Vrelo Vruljak, paratype (SBMNH 632724). B. P. pseudocostellina sp. nov., Blagaj, side spring of Buna at left bank, paratype (SBMNH 626414). C. P. mostarensis Kuščer, 1933, Blagaj Izvor Bune (SBMNH 626411). D. P. mostarensis Kuščer, 1933, Hodbina, Izvor Bunice (SBMNH 626405). Scale bars = 1 mm (SEM SBMNH Vanessa Delnavaz).

Fig. 2 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 2. Photos of studied localities in Hercegovina. A. Konjic. Ljuta spring (LT of Plagigeyeria ljutaensis sp. nov.). B. Blagaj, spring Buna (P. mostarensis var.). C. Konjic, spring at left bank of Ljuta Rivulet (LT of P. konjicensis sp. nov.). D. Donja Jablanica, Komadinovo Vrelo (LT of P. erossi sp. nov.). E. Vučija, Sušicka Jama (P. cf. lewarnei sp. nov.). F. Trebinje, Gorica, spring Vruljak 1 (P. robusta and P. lewarnei sp. nov.). Photos: Jozef Grego, Dorottya Angyal and Mário Olšavský.