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FIG. 10 in Late Triassic to Early Jurassic radiolarian, conodont and ammonite assemblages from the Tavuscayiri block, Mersin Mélange, southern Turkey: Time constraints for the T/J boundary and sedimentary evolution of the southern margin of the northern Neotethys

FIG. 10. — Photomicrographs of the radiolarians from the Orbuklukeli section: A, Palaeosaturnalis subovalis Kozur & Mostler, 1990, Orbuk-44; B, C, Mesosaturnalis artus (Donofrio & Mostler, 1978), Orbuk-21; D, E, Mesosaturnalis octospinus Sugiyama, 1997, Orbuk-21; F, G, Praehexasaturnalis merici Tekin, 2002; F, Orbuk-46; G, Orbuk-47; H, I, Praehexasaturnalis poultoni Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-31; J, K, Praehexasaturnalis tenuispinosus (Donofrio & Mostler, 1978); J, Orbuk-26; K, Orbuk-31; L, M, Praehexasaturnalis tetraradiatus Kozur & Mostler, 1990; L, Orbuk-31; M, Orbuk-32; N-O, Stauroacanthocircus dickinsoni (Yeh, 1989); N, Orbuk-31; O, Orbuk-47; P, Q, Stauroacanthocircus? poetschensis Kozur & Mostler, 1990, Orbuk-31; R, Stauroacanthocircus sp. A, Orbuk-32; S, T, Pseudoacanthocircus mediospinosus Kozur & Mostler, 1990; S, Orbuk-38; T, Orbuk-44; U, Pseudoacanthocircus mocki Kozur & Mostler, 1990, Orbuk-44; V, Pseudoacanthocircus troegeri Kozur & Mostler, 1990, Orbuk-44; W, Pseudoacanthocircus sp. B sensu Sugiyama (1997), Orbuk-31; X, Y, Droltus eurasiaticus Kozur & Mostler, 1990, Orbuk-32. Scale bar: A, 220 µm; B, C, 120 µm; D-G, L-P, R-V, 200 µm; H-K, W, 170 µm; Q, 240 µm; X, Y, 90 µm.

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FIG. 15 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 15. — Viperinae ('Oriental vipers' group) from the early Miocene (MN 4) of MWQ. Middle trunk vertebra Pal. 1994 (2/2003 Reptile Joint) in right lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Posterior trunk vertebra Pal. 1506 (1/2001 Turtle Joint) with an indication of bifurcation in the distal tip of the hypapophysis in left lateral (F), dorsal (G), ventral (H), cranial (I) and caudal (J) views. Abbreviations: cd, condyle; ct, cotyle; d, diapophysis; hy, hypapophysis; na, neural arch; nc, neural canal; pctf, paracotylar foramen; pof, postzygapophyseal articular facet; pp, parapophyseal process; pr, prezygapophysis; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scf, subcentral foramen; scr, subcentral ridge; sctt, subcotylar tubercle; zy, zygosphene; zyg, zygantrum. Scale bars: 5 mm.

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FIG. 13 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 13. — Vipera sp. ('European vipers' group) from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 2022 (2/2003 Reptile Joint) in right lateral (A), dorsal (B), ventral (C) and cranial (D) views. Trunk vertebra Pal. 2023 (2/2003 Reptile Joint) in left lateral (E), dorsal (F), cranial (G) and caudal (H) views. Abbreviations: cd, condyle; ct, cotyle; hy, hypapophysis; nc, neural canal; ns, neural spine; pctf, paracotylar foramen; pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyg, zygantrum. Scale bars: 2 mm.

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FIG. 10 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 10. — "Colubrinae" indet., type 1 from the early Miocene (MN 4) of MWQ. Middle trunk vertebra Pal. 1464 (1/2001 Turtle Joint) in right lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Middle trunk vertebra Pal. 1465 (1/2001 Turtle Joint) in right lateral (F), dorsal (G), ventral (H), cranial (I) and caudal (J) views. Posterior trunk vertebra Pal. 1466 (1/2001 Turtle Joint) in left lateral (K) and ventral (L) views. Caudal vertebra Pal. 1966 (2/2003 Reptile Joint) in right lateral (M), dorsal (N), ventral (O) and cranial (P) views. Abbreviations: cd, condyle; ct, cotyle; d, diapophysis; es, epizypapophyseal spine; hae, haemapophysis; hk, haemal keel; lf, lateral foramen; na, neural arch; nc, neural canal; ns, neural spine; p, parapophysis; pctf, paracotylar foramen; pl, pleurapophysis; pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyg, zygantrum. Scale bar: 2 mm.

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FIG. 12 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 12. — Natrix sp. and "Natricinae" indet. from the early Miocene (MN 4) of MWQ. Natrix sp.: trunk vertebra Pal. 1968 (2/2003 Reptile Joint) in left lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. "Natricinae" indet.: trunk vertebra Pal. 1483 (1/2001 Turtle Joint) with preserved hypapophysis in left lateral (F), dorsal (G), ventral (H) and caudal (I) views. Abbreviations: ak, anterior keel; cd, condyle; ct, cotyle; es, epizypapophyseal spine; hy, hypapophysis; lf, lateral foramen;nc, neural canal; ns, neural spine; pctf, paracotylar foramen; po, postzygapophysis;pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyg, zygantrum. Scale bars: 2 mm.

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FIG. 9 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 9. — Coluber (s.l.) sp., from the early Miocene (MN 4) of MWQ. Anterior trunk vertebra Pal. 1454 (1/2001 Turtle Joint) in right lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Middle trunk vertebra Pal. 1456 (1/2001 Turtle Joint) in left lateral (F), dorsal (G), ventral (H), cranial (I) and caudal (J) views. Anterior caudal vertebra Pal. 1462 (1/2001 Turtle Joint) in right lateral (K), dorsal (L), ventral (M), cranial (N) and caudal (O) views. Abbreviations: cd, condyle; ct, cotyle; d, diapophysis; hae, haemapophysis; hk, haemal keel; hy, hypapophysis; lf, lateral foramen; na, neural arch; nc, neural canal; ns, neural spine; p, parapophysis; pctf, paracotylar foramen; pl, pleurapophysis; pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyg, zygantrum. Scale bars: 2 mm.

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FIG. 8 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 8. — Python sp. from the early Miocene (MN 4) of MWQ. Posterior trunk vertebra Pal. 1449 (1/2001 Turtle Joint) in right lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Abbreviations: cd, condyle; ct, cotyle; hk, haemal keel; na, neural arch; nc, neural canal; ns, neural spine; pd, paradiapophysis; po, postzygapophysis; pof, post zygapophyseal articular facet; pr, prezygapophysis; prf, prezygapophyseal articular facet; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyf, zygosphenal facet; zyg, zygantrum. Scale bar: 5 mm.

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FIG. 7 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 7. — Bavarioboa cf. hermi from the early Miocene (MN 4) of MWQ. Posterior trunk vertebra Pal. 1448 (1/2001 Turtle Joint) in left lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Abbreviations: ct, cotyle; hk, haemal keel; ir, interzygapophyseal ridge; na, neural arch; nc, neural canal; ns, neural spine; pd, paradiapophysis; pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; zy, zygosphene; zyg, zygantrum. Scale bar: 2 mm.

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FIG. 6 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 6. — Ophisaurus and Anguinae indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1407 (1/2001 Turtle Joint) of Ophisaurus sp. in (A) lateral, (B) dorsal, (C) ventral, (D) anterior and (E) posterior aspects. Anguinae indet.: right coronoid Pal. 1408 (1/2001 Turtle Joint) in (F) lateral, (G) medial and (H) dorsal aspects. Osteoderms Pal. 1577-1579 (2/2003 Reptile Joint) in (I-K) external aspects. Abbreviations: alp, anterolateral process; amp, anteromedial process; cd, condyle; ct, cotyle; dp, dorsal process; mr, medial ridge; muc, muscular crest; nc, neural canal; ns, neural spine; nt, notch; os, overlap surface; pof, postzygapophyseal articular facet; pp, posterior process; prf, prezygapophyseal articular facet; rd, ridge; scs, scupltured surface; syn, synapophysis. Scale bars: 2 mm.

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FIG. 11 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 11. — Colubridae gen. et sp. indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1484 (1/2001 Turtle Joint) in right lateral (A), dorsal (B) and cranial (C) views. Abbreviations: ct, cotyle; d, diapophysis; lf, lateral foramen; na, neural arch; nc, neural canal; ns, neural spine; pctf, paracotylar foramen; prf, prezygapophyseal articular facet; zy, zygosphene. Scale bar: 2 mm.

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FIG. 3 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 3. — Amphisbaenia indet. from the early Miocene (MN 4) of MWQ. The trunk vertebra Pal. 1570 (2/2003 Reptile Joint) in (A) lateral, (B) dorsal, (C) ventral, (D) anterior and (E) posterior aspects. Abbreviations: cd, condyle; ct, cotyle; nc, neural canal; po, postzygapophysis; pof, postzygapophyseal articular fac- et; pr, prezygapophysis; prf, prezygapophyseal articular facet; scf, subcentral foramen; syn, synapophysis. Scale bar: 1 mm.

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FIG. 16 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 16. — Elapidae gen. et sp. indet. from the early Miocene (MN 4) of MWQ. Trunk vertebra Pal. 1500 (1/2001 Turtle Joint) in right lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views. Abbreviations: cd, condyle; ct, cotyle; hy, hypapophysis; na, neural arch; nc, neural canal; ns, neural spine; pctf, paracotylar foramen; pof, postzygapophyseal articular facet; prf, prezygapophyseal articular facet; scg, subcentral groove; scr, subcentral ridge; sctt, subcotylar tubercle; zyg, zygantrum. Scale bar: 2 mm.

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FIG. 14 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 14. — Viperinae ('Oriental vipers' group) from the early Miocene (MN 4) of MWQ. Left maxilla Pal. 1501 (1/2001 Turtle Joint) in rostral (A) and caudal (B) views. Fang Pal. 1502 (1/2001 Turtle Joint) in anterolateral (C) view. Anterior trunk vertebra Pal. 1503 (1/2001 Turtle Joint) in right lateral (D), dorsal (E), ventral (F) and cranial (G) views. Trunk vertebra Pal. 1504 (1/2001 Turtle Joint) in right lateral (H) and cranial (I) views. Vertebra from the middle part of the trunk section Pal. 1505 (1/2001 Turtle Joint) in right lateral (J), dorsal (K), ventral (L), cranial (M) and caudal (N) views. Abbreviations: can.den, dental canal; cd, condyle; ct, cotyle; fac. ecpt, facet for contact with ectopterygoid; hy, hypapophysis; ir, interzygapophyseal ridge; na, neural arch; nc, neural canal; ns, neural spine; pctf, paracotylar foramen; pd, paradiapophysis; po, postzygapophysis; pof, postzygapophyseal articular facet; pp, parapophyseal process; pr, prezygapophysis; pr.asc, ascending process; prf, prezygapophyseal articular facet; r, ridge; scf, subcentral foramen; scg, subcentral groove; scr, subcentral ridge; sctc, subcotylar canal; zy, zygosphene; zyg, zygantrum. Scale bars: A-C, 2 mm; D-N, 5 mm.

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FIG. 2 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 2. — Lacertidae indet. from the early Miocene (MN 4) of MWQ. Left frontal Pal. 1565 (2/2003 Reptile Joint) in (A) dorsal and (B) ventral aspects. Lacertidae indet. tooth morphotype 1: right maxilla Pal. 1566 (2/2003 Reptile Joint) in (C) lateral and (D) medial aspects with detail of teeth. Left dentary Pal. 1567 (2/2003 Reptile Joint) in (E) lateral and (F) medial aspects with detail of teeth. Right dentary Pal. 1568 (2/2003 Reptile Joint) in (G) medial aspect with detail of teeth. Lacertidae indet. tooth morphotype 2: left dentary Pal. 1401 (1/2001 Turtle Joint) in (H) medial aspect with detail of teeth. Abbreviations: dc, dental crest; fs, frontal shield; lf, labial foramen (foramina); lp, lateral process; mc, mesial cusp; Mg, Meckel`s groove; mp, medial process; prs, prefrontal shield; sbs, subdental shelf; sc, sulcus; srs, supradental shelf; st, striations. Scale bars: A-D, H, 1 mm; D', G', 100 µm; E-G, H', 500 µm; F', 200 µm.

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FIG. 1 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 1. — Topographic position of the Mokrá-Western Quarry with 1/2001 Turtle Joint and 2/2003 Reptile Joint (A). Position of the 1/2001 Turtle Joint at floor 380 m a.s.l. and field excavation of the fissure deposits in 2002 (B). The 2/2003 Reptile Joint has been discovered only few meters on the left from the 1/2001 Turtle Joint (modified after Ivanov et al. 2006).

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FIG. 5 in Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications

FIG. 5. — Pseudopus from the early Miocene (MN 4) of MWQ. Pseudopus laurillardi: parietal Pal. 1574 (2/2003 Reptile Joint) in (A) dorsal and (B) ventral aspects; parietal Pal. 1402 (1/2001 Turtle Joint) in (C) dorsal and (D) ventral aspects. Pseudopus sp.: left maxilla Pal. 1575 (2/2003 Reptile Joint) in (E) lateral and (F) medial aspects. Left quadrate Pal. 1576 (2/2003 Reptile Joint) in (G) lateral and (H) medial aspects. Right dentary Pal. 1403 (1/2001 Turtle Joint) in (I) lateral, (J) medial and (K) dorsal aspects. Trunk vertebra Pal. 1404 (1/2001 Turtle Joint) in (L) lateral, (M) dorsal, (N) ventral, (O) anterior and (P) posterior aspects. Trunk vertebra 1405 (1/2001 Turtle Joint) in (Q) anterior aspect. Abbreviations: are, arcuate edge; cc, cephalic condyle; cd, condyle; cf, facet for

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aquila (Whalen & Carter in Carter, Whalen & Guex, 1998), Orbuk-32; R, Pseudoeucyrtis busuangaensis (Yeh & Cheng, 1998), Orbuk-32; S, Farcus graylockensis Pessagno, Whalen & Yeh, 1986, Orbuk-32; T-U, Farcus sp. A, Orbuk-26; V, Farcus sp. B, Orbuk-27; W-X, Anaticapitula anatiformis (De Wever, 1982); W, Orbuk-32; X, Orbuk-38; Y, Saitoum sp. aff. S. triumphense Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-32. Scale bar: A-C, K-L, T-U, W-Y, 100 µm; D-J, M-O, 120 µm; P-S, V, 150 µm. in Late Triassic to Early Jurassic radiolarian, conodont and ammonite assemblages from the Tavuscayiri block, Mersin Mélange, southern Turkey: Time constraints for the T/J boundary and sedimentary evolution of the southern margin of the northern Neotethys

aquila (Whalen & Carter in Carter, Whalen & Guex, 1998), Orbuk-32; R, Pseudoeucyrtis busuangaensis (Yeh & Cheng, 1998), Orbuk-32; S, Farcus graylockensis Pessagno, Whalen & Yeh, 1986, Orbuk-32; T-U, Farcus sp. A, Orbuk-26; V, Farcus sp. B, Orbuk-27; W-X, Anaticapitula anatiformis (De Wever, 1982); W, Orbuk-32; X, Orbuk-38; Y, Saitoum sp. aff. S. triumphense Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-32. Scale bar: A-C, K-L, T-U, W-Y, 100 µm; D-J, M-O, 120 µm; P-S, V, 150 µm.

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FIG. 7 in Late Triassic to Early Jurassic radiolarian, conodont and ammonite assemblages from the Tavuscayiri block, Mersin Mélange, southern Turkey: Time constraints for the T/J boundary and sedimentary evolution of the southern margin of the northern Neotethys

FIG. 7. — Photomicrographs of the radiolarians from the Orbuklukeli section: A-D, Praeudalia rhaetica Tekin, n. gen., n. sp.; A, Holotype, Orbuk-15; B-D, Paratypes, Orbuk-15; E, Udalia dennisoni Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-32; F-G, Udalia infrequens Tekin, n. sp.; F, Holotype, Orbuk-36; G, Paratype, Orbuk-36; H-J, Udalia primaeva Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-27; K-M, Thurstonia gibsoni Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-26; N, Thurstonia minutaglobus Whalen & Carter in Carter, Whalen & Guex, 1998, Orbuk-32; O, Thurstonia timberensis Whalen & Carter in

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FIG. 6 in Late Triassic to Early Jurassic radiolarian, conodont and ammonite assemblages from the Tavuscayiri block, Mersin Mélange, southern Turkey: Time constraints for the T/J boundary and sedimentary evolution of the southern margin of the northern Neotethys

FIG. 6. — Photomicrographs of the radiolarians from the Orbuklukeli section: A, Betraccium kennecottense Carter, 1993, Orbuk-15; B, Betraccium perilense Carter, 1993, Orbuk-15; C, D, Gorgansium alpinum Kozur & Mostler, 1990, Orbuk-32; E, F, Gorgansium gongyloideum Kishida & Hisada, 1985, Orbuk-41; G, Pantanellium fosteri Pessagno & Blome, 1980, Orbuk-15; H-J, Pantanellium freboldi Whalen & Carter in Carter, Whalen & Guex, 1998; H-I, Orbuk-27; J, Orbuk-31; K-O, Pantanellium giganteum Tekin, n. sp.; K, Holotype, Orbuk-26; L-O, Paratypes; L, Orbuk-26; M, Orbuk-27; N-O, Orbuk-31; P, Pantanellium kluense Pessagno & Blome, 1980, Orbuk-26; Q, R, Pantanellium tanuense Pessagno & Blome,1980; Q, Orbuk-26; R, Orbuk-27; S-U, Novamuria impensa (Whalen & Carter in

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FIG. 12 in Late Triassic to Early Jurassic radiolarian, conodont and ammonite assemblages from the Tavuscayiri block, Mersin Mélange, southern Turkey: Time constraints for the T/J boundary and sedimentary evolution of the southern margin of the northern Neotethys

FIG. 12. — Photomicrographs of the radiolarians from the Orbuklukeli section; A, B, Canoptum rhaeticum Kozur & Mostler, 1981, Orbuk-15; C, Canoptum striatum (Kozur & Mostler, 1990), Orbuk-27; D-J, Laxtorum breve Tekin, n. sp.; D, Holotype, Orbuk-32; E-J, Paratypes, Orbuk-32; K-L, Laxtorum obscurum Tekin, n. sp., K, Holotype, Orbuk-27; L, Paratype, Orbuk-27; M-N, Atalantria emmela (Cordey & Carter, 1996), Orbuk-32; O, Atalantria sp. A, Orbuk-27; P-Q, Pseudoeucyrtis

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Last verified 2026-04-29Open record