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Fig. 7 in Moulladella Jourdanensis (Foury & Moullade, 1966) N. Gen., N. Comb.: Valanginian-Early Late Barremian Larger Benthic Foraminifera From The Northern Neotethyan Margin

Fig. 7 Benthic foraminifera associated with Moulladella jourdanensis (Foury & Moullade) n. gen., n. comb. from the Dâmbovicioara area, Romania. (a) Coscinoconus campanellus (Arnaud-Vanneau, Boisseau & Darsac), slightly oblique axial section. (b) Coscinoconus cherchiae (Arnaud-Vannaeu, Boisseau & Darsac), axial section. (c) Danubiella gracillima Neagu, subaxal section. (d) Protopeneroplis ultragranulata (Gorbatchik), oblique section. (e, h) Paracoskinolina? pfenderae Canerot & Moullade, axial and slightly oblique transverse sections. (f, g) Montsalevia salevensis (Charollais, Brönnimann & Zaninetti), slightly oblique axial section, and transverse section together with Haplophragmoides joukowsky Charollais, Brönnimann & Zaninetti (axial section) (H). Thin-sections: 14096 (a), 12562 (b), 12569 (c), 13697 (d), 12568- 2 (e), 12563 (f), 12564 (g) and 11867-6 (h). Scale bars 0.3 mm.

Fig. 10 Persiella pseudolituus n. gen., n in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 10 Persiella pseudolituus n. gen., n. sp. from the upper Maastrichtian of the Tarbur Formation of the Naghan (a) and Mandegan (all others) sections, Zagros Zone, SW Iran. a–c, h Oblique equatorial sections. d, f–g Equatorial sections. e Equatorial section of an uncoiling specimen. Thin-sections: NG 197-2 (a), Rt 72 (b), Rt 79 (c), Rt 113 (d, f–h), Rt 98 (e).

Fig. 12 Persiella pseudolituus n. gen., n in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 12 Persiella pseudolituus n. gen., n. sp. from the upper Maastrichtian of the Tarbur Formation of the Mandegan (a, g, i–j) and Naghan sections (all others), Zagros Zone, SW Iran. a–e, h–i Oblique sections; note the rounded outline of the network in a and b. f Tangential section; note the rounded subepidermal network (lower part). g Almost centered axial section, holotype specimen. j Juvenile specimen showing protoconch connected to the following chamber (?deuteroconch) by numerous openings. Abbreviations: b = beam, p = proloculus, d? = possible deuteroconch, f = foramen, r = rafter, s = septum. Thin-sections: Rt 87 (a), NG 198 (b), 2NG 120-1 (c), 2NG 144 (d), 2NG 113 (e), 2NG 200-1 (f), Rt 113 (SNSB-BSPG 2016 V 16) (g), 2NG 200-1 (h), Rt 113-1 (i), Rt 72 (j).

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Fig. 4 in Moulladella Jourdanensis (Foury & Moullade, 1966) N. Gen., N. Comb.: Valanginian-Early Late Barremian Larger Benthic Foraminifera From The Northern Neotethyan Margin

Fig. 4 Moulladella jourdanensis (Foury & Moullade) n. gen., n. comb. (a, g) Tangential-oblique sections of microspheric specimens exhibiting voluminous initial spire. (b) Axial section of a macrospheric specimen. (c, k) Tangential sections showing alternating chamberlets of the marginal zone connected by oblique foramina. (d) Oblique-tangential section. (e, j, m) Axial sections. (h, l) Oblique sections. (f, i) Juvenile specimens in tangential sections. (n) Oblique section of the initial spire of a macrospheric specimen. b, beam; f, foramen; p, protoconch; pi, pillar, s, septum. Thin-sections: 14729 (a), 12563-5 (b), 14099-2 (c), 12561-2 (d, f), 12558A (e), 42-Irimieş (g), 12562-5 (h), 12563-4 (i), 12563-6 (j), 14098-2 (k), 12563 (l), 12563-2 (m), 12563-4 (n). Location and age: (a) Jerma River Canyon, Eastern Serbia, Early Valanginian; (b–f, h–n) Dâmbovicioara area, Southern Carpathians, Romania (Early Valanginian); (g) Reşiţa-Moldova Nouă zone, Southern Carpathians, Romania (Early Barremian).

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Fig. 3 in Moulladella Jourdanensis (Foury & Moullade, 1966) N. Gen., N. Comb.: Valanginian-Early Late Barremian Larger Benthic Foraminifera From The Northern Neotethyan Margin

Fig. 3 Paracoskinolina (Meyendorffina) jourdanensis Foury & Moullade, Barremian of France (a–d); holotype specimen in (a) (from Foury and Moullade, 1966, pl. 1, figs. 2–3, 5–6). e–h Conicopfenderina? balkanica Peybernès, Valanginian of Bulgaria; holotype specimen in (f) (from Peybernès, 2004, pl. 1, figs. 7, 1, 8–9).

Fig. 9 Persiella pseudolituus n. gen., n in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 9 Persiella pseudolituus n. gen., n. sp. from the upper Maastrichtian of the Tarbur Formation of the Mandegan (a, e, g) and Naghan sections (all others), Zagros Zone, SW Iran. a–b Equatorial sections. c–d, g Oblique equatorial sections; note tendency to uncoil in the youngest chambers. e–f Oblique equatorial sections of uncoiling specimens. Abbreviations: p = proloculus, d = possible deuteroconch, f = foramen, s = septum. Thin-sections: Rt 113 (a, g), 2NG 191 (b), 2NG 186 (c), 2NG 49 (d), Rt 87-1 (e), NG 196 (f).

Fig. 8 in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 8 Schematic drawing showing differences in the wall structure of Persiella n. gen. and Pseudocyclammina Yokoyama (without scale).

Fig. 7 Flabelloperforata tarburensis n. gen., n in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 7 Flabelloperforata tarburensis n. gen., n. sp. from the upper Maastrichtian of the Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a Oblique section showing multiple foramina. b Detail from a showing pseudo-keriotheka-like wall structure. c Oblique section. d, e Details (d from c) showing multiple foramina. f Oblique section. Thin-sections: 2NG 8 (a–b), 2NG 86 (c–e), 2NGN (f). Scale bars: a–b = 0.2 mm, c, f = 1 mm, d–e = 0.3 mm.

Fig. 6 Flabelloperforata tarburensis n. gen., n in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 6 Flabelloperforata tarburensis n. gen., n. sp. from the upper Maastrichtian of the Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a–e, g–l: Oblique sections; holotype specimen in a; multiple foramina depicted in i and k with arrows. f Oblique equatorial section. T: Tarburina zagrosiana Schlagintweit, Rashidi & Barani. m Slightly oblique subaxial section. Arrow: multiple foramina. Thin-sections: NG 85-2 (a), 2NG 81-2 (b), 2NG 85-1 (c), 2NG 83 (d), 2NG 84-1 (e, h), NG 49 (f), 2NG 81-4 (g), NG 83-2 (i), NG 81 (j), NG 61 (k), 2NG 87-2 (l), 2NG 197 (m).

Fig. 1 in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 1 Lithostratigraphy of Upper Cretaceous strata of the Zagros Zone (redrawn from the Stratigraphic Chart of Iran, Geological Society of Iran, 1995).

Fig. 5 in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 5 Neobalkhania bignoti Cherchi, Radoičić & Schroeder from the upper Maastrichtian of the Tarbur Formation of the Mandegan (a), and Naghan sections (all others). a Subaxial section of a large specimen (diameter ~7 mm). b Axial section of a megalospheric specimen. c Four specimens in (sub)axial sections, and Dicyclina sp. (D). d Tangential section showing subepidermal network with coarse primary partitions (right below) and finer secondary partitions (left below). e Oblique axial section of a megalospheric specimen. f Fragmentary equatorial section, P = Persiacyclammina maastrichtiana Schlagintweit & Rashidi. g Oblique section of a megalospheric specimen showing large embryo with opening to the second annular chamber. Scale bars: a–b, d, f–g = 0.3 mm; C, E = 0.6 mm. Thin-sections: Rt 113 (a), 2NG 109 (b), 2NG 128 (c), 2NG 201-1 (d), 2NG 120 (e), 2NG 81 (f–g).

Fig. 3 in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 3 Typical microfacies of samples with Flabelloperforata tarburensis n. gen., n. sp (F in a–b), and Persiella pseudolituus n. gen., n. sp. (Ps in c–d), from the upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a–b Wackestones/packstones with benthic foraminifera, among many porcelaneous and agglutinating taxa such as Dicyclina sp. (Di), Tarburina zagrosiana Schlagintweit, Rashidi & Barani (T), Dictyoconus bakhtiari Schlagintweit, Rashidi & Babadipour (D) and remains of dasycladalean algae (d in b). c Wackestone with small and larger benthic foraminifera and oblique longitudinal section of the dasycladale Pseudocymopolia cf. anadyomenea (Elliott) (P). d Grainstone/packstone with Omphalocyclus macroporus Lamarck (O), Dictyoconella? minima Henson (Dic), and slightly oblique axial section of Persiella pseudolituus n. gen., n. sp. (Ps). Thin-sections: 2NG 81-4 (a), 2NG 120 (b), 2NG 166 (c), 2NG 177 (d).

Fig. 2 in Persiella Pseudolituus N. Gen., N. Sp., And Flabelloperforata Tarburensis N. Gen., N. Sp., Two New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran

Fig. 2 Location of the Naghan and Mandegan sections, the type localities of Flabelloperforata tarburensis n. sp., and Persiella pseudolituus n. gen., n. sp. (taken from google maps)

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Fig. 3 in Persiacyclammina Maastrichtiana N. Gen., N. Sp., A New Larger Benthic Foraminifer From The Maastrichtian Of Iran

Fig. 3 Wall-structure of Persiacyclammina maastrichtiana n. gen., n. sp. from the upper Maastrichtian of SW Iran Thin-section: NG 83-1.

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Fig. 1 in Persiacyclammina Maastrichtiana N. Gen., N. Sp., A New Larger Benthic Foraminifer From The Maastrichtian Of Iran

Fig. 1 Location of the Naghan section, the type locality of Persiacyclammina maastrichtiana n. gen., n. sp.

Fig. 6 Zagrosella rigaudii n.gen., n in Zagrosella Rigaudii N. Gen., N. Sp., A New Biokovinoidean Foraminifer From The Maastrichtian Of Iran

Fig. 6 Zagrosella rigaudii n.gen., n.sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a– d, g, Oblique equatorial sections. e–f, h–i, Oblique sections. Abbreviations: p = proloculus, pi = pillar, t = structures interpreted as cryptoendolithic thaumatoporellaceans. Thin-sections: 2NG 98 (a), NG 92 (b), NG 91 (c), 2NG 85 (d), NG 49 (e), NG 86 (f), 2NG 84 (g), NG 83-2 (h), NG 84 (i)

Fig. 5 Zagrosella rigaudii n.gen., n in Zagrosella Rigaudii N. Gen., N. Sp., A New Biokovinoidean Foraminifer From The Maastrichtian Of Iran

Fig. 5 Zagrosella rigaudii n.gen., n.sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, c, Equatorial sections, b, d–g, Oblique equatorial sections. h, Equatorial section of a microspheric specimen. Abbreviations: p = proloculus, pi = pillar, t = structures interpreted as cryptoendolithic thaumatoporellaceans. Thin-sections: NG 88 (a), 2NG 94 (b), 2NG 91 (c), 2NG 85 (d), NG 88 (e), NG 14 (f), 2NG 98 (g), NG 38-1 (h).

Fig. 4 in Zagrosella Rigaudii N. Gen., N. Sp., A New Biokovinoidean Foraminifer From The Maastrichtian Of Iran

Fig. 4 Wall-structure of Zagrosella rigaudii n.gen., n.sp. from the upper Maastrichtian of SW Iran (a–b) and Biokovina gradacensis Gušić from the Liassic of Croatia (c–d). Polygonal parapores are marked with an arrow in a. Thin-sections: 2NG 27 (a), 2NG 87 (b). Remarks and comparisons: The wall structure of Zag- the Late Cretaceous, it can be compared to some extent rosella with its "uniform, parallel, radial elements cov- (e.g., similar wall-structure, strengthenings) with Braciered by some kind of tectum" (Hottinger, 2006, p. 29) can ana Schlagintweit & Cvetko-Tesovic, 2016 (Santonian?- be compared with the keriotheca-like (or pseudokerithe- lower Campanian of Croatia). The latter differs from cal) texture of some Mesozoic larger benthic foraminifera Zagrosella n.gen. above all by its test morphology (elon- (e.g., Schroeder et al., 1975; De Castro, 1981; Septfon- gate compressed), and the foraminal characteristics (sevtaine, 1981; Banner et al., 1991; Rigaud et al., 2015). eral close-set openings, equal in diameter to the wall al- Regarding differences to parapores (or canaliculi), the veolae, and may have short peristomal rims at their mardifferences are vague and not clearly delimited (Hot- gins) (see Schlagintweit and Cvetko-Tešović, 2016 for tinger, 2006: "usually much larger"…"often more irregu- details). lar"). Zagrosella n.gen. is morphologically very similar to Last but not least we note morphological similarities the Liassic Biokovina Gušić, 1977, both monospecific to the Late Jurassic Labyrinthina Weynschenk that differs genera. The wall structure of Zagrosella appears more above all from Zagrosiella by its wall being "agglutinatevolved with larger parapores (diameter 0.01 to 0.03 mm, ed, simple in structure, microgranular, imperforate" see description below) whereas in Biokovina these are (Loeblich and Tappan (1987, p. 96). thinner (but with overlapping range), and bifurcating (diameter ~0.01 to 0.15 mm). There are also differences in Zagrosella rigaudii Schlagintweit & Rashidi, n.sp. the endoskeletal structures, being more massive (in the Figs. 3 pars, 4a–b, 5–7 central part of the test) and may fuse in Biokovina where- Origin of the name: The species name refers to Sylvain as in Zagrosella they occur preferentially on the sides of Rigaud (Singapore) for his outstanding contributions to the chambers, sometimes only partially differentiated the micropalaeontology and phylogeny of benthic foramfrom the chamber wall, and never fused. They are here inifera. interpreted as pillars (originating from the septa), not as Holotype: Slightly oblique equatorial section illustrated septula (originating from the wall) (see Hottinger, 2006) in Figure 5A, thin-section NG 88. or strengthenings (see Rigaud et al., 2013). Another mor- Paratypes: Figs. 5B–G, 6, 7A–B. phologically similar genus of the family Biokovinidae is Description: Test free, lenticular, with oscillating plane Bosniella Gušić, 1977. It differs from Zagrosella above of coiling in the early stage, later planispiral, and finally all by its chambers lacking any endoskeletal structures. In maybe uncoiling, rectilinear. It is compressed axially,

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Fig. 8 Persiacyclammina maastrichtiana n. gen., n in Persiacyclammina Maastrichtiana N. Gen., N. Sp., A New Larger Benthic Foraminifer From The Maastrichtian Of Iran

Fig. 8 Persiacyclammina maastrichtiana n. gen., n. sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, f–h, l, Oblique sections. b–d, Oblique sections through the uncoiled part. i–j, Transverse sections, slightly oblique of the uncoiled part. k, subtransverse section. m–n, Details of oblique sections showing structural elements. Abbreviations: a.l. = alveolar layer, f = foramen, r = rafter, s = septum, s.s. = subepidermal septulum, r.n. = reticulate network. Thin-sections: 2NG 81-2 (a), 2NG 55 (b), 2NG 114 (c), 2NG 1 (d), NG 74 (e), 2NG 81-4 (f, h), 2NG 81-1 (g), 2NG 71 (i), NG 83-3 (j), NG 81 (k), 2NG 18-2 (l), 2NG 85-4 (m), 2NG 81-3 (n).

Fig. 8 in Zagrosella Rigaudii N. Gen., N. Sp., A New Biokovinoidean Foraminifer From The Maastrichtian Of Iran

Fig. 8 Biokovina gradacensis Gušić, Liassic of Croatia. Photographs made from the original thin-section material of Gušić (1977: A see plate 2, fig. 2, and B see plate 1, fig. 2).