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FIGURE 21 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale.

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FIGURE 20 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika.

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FIGURE 22 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 22. Photos of Lygodactylus fritzi sp. nov. specimens from Vohibola in life. All specimens correspond to genotyped individuals included in Fig. 1 that were not collected.

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FIGURE 19 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 19. Photo of Lygodactylus hapei sp. nov., holotype ZSM 298/2018 from Djangoa (Djohahely), in life. Photo by H.-P. Berghof.

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FIGURE 18 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 18. Photos of Lygodactylus roellae sp. nov. in life. A and B, adult female holotype ZSM 49/2016 (MSZC 0072) from Ampotsidy. C,D, adult female paratype UADBA-R 70855 from Ampotsidy. E, male paratype ZSM 556/2014 (DRV 6289) from Andrevorevo.

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FIGURE 16 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 16. Photos of specimens of Lygodactylus tantsaha sp. nov. from Montagne d'Ambre in life. A, Adult male holotype, ZSM 196/2018 (MSZC 0772); B, specimen MSTIS 928 (voucher not collected); C, D, Adult male paratype, ZSM 197/2018 (MSZC 0771). Not to scale.

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FIGURE 17 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 17. Photo of Lygodactylus salvi sp. nov., adult male holotype ZSM 783/2001 (FGMV 2001.74) from Tsaratanana (Antsahamanara) in life.

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FIGURE 15 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study.

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FIGURE 13 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 13. Lygodactylus miops in life. A, specimen from Nahampoana probably assignable to this species, photographed 1992 (not genotyped and not assignable to a voucher specimen). B–F, specimens from Betampona, all genetically identified by DNA barcoding with sequences of the COI gene (voucher specimens not collected). B, sample number ACZC 5544 (ACP1884); C, sample number ACZC 5666 (ACP1947); D, sample number ACZC 5478 (ACP1843); E, sample number ACZC 5420 (ACP1807); F, sample number ACZC 5467 (ACP1839).

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FIGURE 12 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 12. Photos of specimens of Lygodactylus petteri from Montagne d'Ambre in life. A, Adult male ZSM 195/2018 (MSZC 485); B, C, Female ZSM 194/2018 (MSZC 454) in dorsolateral and ventral views. Not to scale.

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FIGURE 14 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 14. Photos of specimens of L. guibei in life. A, female specimen ZSM 525/2009 (ZCMV 11210) from the western slope of the Makira Massif, photographed on 23 June 2009. B–C, male and D–E, female, from Andasibe, in dorsolateral and ventral views, photographed 2003, not attributable to a voucher specimen and not genotyped, and thus only tentatively attributable to the species.

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FIGURE 8 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 8. Photos of specimens of Lygodactylus expectatus from Ankarana in life. A, B, Adult male ZSM 284/2004 (FGZC 543), photographed in 2004. C, D, E, Specimen photographed in 2001 by Gerardo García, not collected.

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FIGURE 11 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 11. Specimens of Lygodactylus madagascariensis in life. A, B, Specimens from the type locality Nosy Be photographed in 1992 (not corresponding to any voucher or sample studied herein; identification by typical color pattern and provenance. C, D, Adult male ZSM 832/2003 from Manongarivo, photographed 2003.

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FIGURE 5 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 5. Tail base in males of different species of the Lygodactylus madagascariensis group, showing presence of distinct tail base tubercles in species primarily belonging to the Domerguella subclade A5 (panels D–K) and its absence in two other species (panels A–C). A–B, dorsal and lateral view of tail base in L. petteri, ZSM 195/2018. C, lateral view of tail base in L. expectatus, ZSM 1540/2008. D–E, dorsal and lateral view of tail base in L. winki sp. nov., ZSM 48/2016. F, lateral view of tail base in L. ulli sp. nov., ZSM 154/2005. G, lateral view of tail base in L. guibei, ZSM 349/2010. H, lateral view of tail base in L. salvi sp. nov., ZSM 783/2001 (subclade A3). I, lateral view of tail base in life of a specimen assigned to L. guibei. J, dorsal view of tail base in life of a specimen from Nahampoana assigned to L. miops. K, ventral view of tail base in life in a specimen assigned to L. guibei. Blue arrows point to lateral tail base tubercles. Whitish structures in A, B, E, F, H are everted hemipenes. All specimens facing to the left. Not to scale.

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FIGURE 2 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 2. Haplotype networks constructed from partial sequences of the nuclear-encoded genes for RAG1 (784 bp) and CMOS (376 bp) for species of the Lygodactylus madagascariensis group. Small blue dots represent additional mutational steps or unsampled haplotypes. Colors match those of allocation of individuals to species-level lineages inferred from the 16S dataset and represented in the mtDNA tree (Fig. 1). The networks were constructed from phased sequences and each individual is therefore included twice in each network (circle size proportional to the total number of haplotype sequences).

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FIGURE 10 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 10. Specimen of Lygodactylus rarus from Ankarana in life, photographed in 2003 (not corresponding to any of the morphologically examined voucher specimens).

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FIGURE 1. Majority-rule consensus tree from a in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 1. Majority-rule consensus tree from a Maximum likelihood analysis based on 537 bp of the mitochondrial 16S rRNA gene, for all available samples of the Lygodactylus madagascariensis group. Outer circles of different color mark categories of bootstrap branch support as indicated, inner circles of different color mark categories of Posterior Probability support from a separate Bayesian analysis. Missing circles on the backbone of clade A5 indicate different topology in the ML and BI analysis. Colors correspond to species-level lineages delimited by the ASAP partition with lowest ASAP score, with two exceptions: (i) L. madagascariensis samples from Montagne d'Ambre are shown as different clusters to better illustrate its co-occurrence with other lineages at this site, as well as patterns of allele sharing in the nuclear encoded genes (Fig. 2); (ii) the three uppermost samples of L. guibei were defined by ASAP as separate lineage but are here considered as conspecific with L. guibei in a preliminary way.

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FIGURE 9 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 9. Tails of preserved specimens of Lygodactylus rarus and six other species of the subgenus Domerguella for comparison. Note the very regular dark-light crossbands characterizing L. rarus, including the holotype. Not to scale.

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FIGURE 7 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 7. Lateral view of head and anterior body in seven specimens of Lygodactylus expectatus, and four representative individuals of other species. In all L. expectatus, a black spot is visible in the shoulder region, with at least traces of light color along its dorsal edge; this spot, poorly expressed in some specimens, is here called the scapular semi-ocellus. Individuals of other species of Domerguella may also have dark markings in the scapular region, but these usually are not at all bordered by light color dorsally, and often are in a more anterior position, as exemplified by four representative pictures of other species. Asterisks mark photos that were mirrored. Pictures not to scale.

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FIGURE 6 in Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar's rainforest

FIGURE 6. View of central dorsum (posterior to forelimb insertion) in seven specimens of Lygodactylus expectatus, and four representative individuals of other species. In all L. expectatus, including the holotype, the scales in the dorsolateral area are distinctly enlarged, while the vertebral scales are much smaller. In the other species, all scales are roughly of similar size, without obviously enlarged dorsolateral scales. Pictures not to scale.

opennotspecifiedAug 2022View details →

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Allen Brain Atlas

Allen Brain Atlas is an Allen Institute collection of brain map atlases, datasets, APIs, and analysis tools covering mouse, human, and non-human primate brain resources.

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neuroscienceopenDocumentation, web resources, and API references are available online.
Last verified 2026-04-30Open record

Annotated Behaviour and Observability Dataset (ABODe)

ABODe is a University of Edinburgh DataShare dataset for behavior classification in group-housed mice using home-cage video, identities, bounding boxes, ground-plate positions, and annotator labels.

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behavioral-neuroscienceopenThe DataShare record exposes download links for annotations, documentation, license text, and the zipped per-snippet data directory.
Last verified 2026-04-30Open record

DANDI Archive for NWB datasets

DANDI is a BRAIN Initiative archive for publishing and sharing neurophysiology data, including electrophysiology, optophysiology, and behavioral data packaged as NWB and related standards.

dandi-nwb
electrophysiologyopenPublished Dandiset metadata and archive endpoints are available through the production DANDI API.
Last verified 2026-04-30Open record

International Brain Laboratory public data

The International Brain Laboratory public data releases expose standardized mouse decision-making experiments, including Neuropixels recordings, widefield calcium imaging, behavior, and session metadata accessed through the ONE API.

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behavioral-neuroscienceopenPublic sessions can be searched and loaded from the IBL public data server through ONE.
Last verified 2026-04-29Open record

OpenNeuro

OpenNeuro is a free, open platform for sharing neuroimaging datasets, with public search, dataset pages, and download paths for web, S3, DataLad, and the OpenNeuro CLI.

openneuro
neuroscienceopenPublished datasets are available on demand over the internet.
Last verified 2026-04-29Open record

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