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FIGURE 19 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 19. Principal Component Analysis (PCA) based on shell ratios and apical angle (SL/MD, AL/AW, SL/LWH, LWH/ AH and AA). A. Scatter of all measured specimens. B. Cluster defining genus groups. Abbreviations: age: Astyris? agenta Harzhauser & Kowalke, 2002, aus: Auingeria austriaca (Hoernes & Auinger, 1880), bel: Auingeria bellardii (Hoernes, 1852), bit: "Mitrella" bittneri (Hoernes & Auinger, 1880), bro: Thiarella bronni (Mayer, 1869), buc: Defensina bucciniformis (Hoernes & Auinger, 1880), com: Mitrella complanata (Sacco, 1890), con: Orthurella convexula Sacco, 1904, dac: Mitrella dacica nov. sp., dem: Mitrella demaintenonae nov. sp., duj: Martaia dujardini (Hoernes & Auinger, 1880), elo: Mitrella elongatissima nov. sp., emb: Bellacolumbella embryonalis (Boettger, 1902), fal: Mitrella fallax (Hoernes & Auinger, 1880), gue: Costoanachis guembeli (Hoernes & Auinger, 1880), gru: Auingeria grussbachensis (Csepreghy-Meznerics, 1969), hau: Scabrella haueri (Hoernes & Auinger, 1880), hil: Mitrella hilberi (Cossmann, 1901), kar: Bellacolumbella karreri (Hoernes & Auinger 1880), kat: Bellacolumbella katharinae (Hoernes & Auinger, 1880), kor: Graphicomassa korytnicensis (Bałuk, 1995), lat: Costoanachis latecostata (Boettger, 1902), may: Columbella mayeri Hoernes, 1856, mor: Orthurella moravica (Hoernes & Auinger, 1880), pet: Mitrella petersi (Hilber, 1879), pol: Graphicomassa polonica (Pusch, 1837), sar: Astyris sarmatica (Kojumdgieva, 1969), sop: Clinurella sophiae Boettger, 1902, sph: Zafrona sphaerocorrugata nov. sp., vie: Mitrella viennensis nov. sp., zit: Martaia zitteli (Hoernes & Auinger, 1880).

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FIGURE 17. A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 17. A1–A2. Bellacolumbella embryonalis (Boettger, 1902), SMF 360676, lectotype, Coşteiu de Sus (Romania). B1–B2. Bellacolumbella embryonalis (Boettger, 1902), SMF 360676, paralectotype, Coşteiu de Sus (Romania). C1–C2. Bellacolumbella karreri (Hoernes & Auinger 1880), NHMW1869/0001/0166b, paralectotype, Baden (Austria), D1–D2. Bellacolumbella karreri (Hoernes & Auinger 1880), NHMW1846/0037/0147, lectotype, Baden (Austria). E. Bellacolumbella karreri (Hoernes & Auinger 1880), NHMW1846/0037/0147a, paralectotype, Baden (Austria): F1–F2. Bellacolumbella karreri (Hoernes & Auinger 1880), NHMW1869/0001/0166a, paralectotype, Baden (Austria). G1–G2. Bellacolumbella katharinae (Hoernes & Auinger, 1880), NHMW1865/0001/0645a, lectotype, Sedlec (the Czech Republic). H1–H2. Bellacolumbella katharinae (Hoernes & Auinger, 1880), NHMW1865/0001/0645b, paralectotype, Sedlec (the Czech Republic). I1–I2. Bellacolumbella katharinae (Hoernes & Auinger, 1880), NHMW1859/0019/0084, paralectotype, Sedlec (the Czech Republic).

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FIGURE 14. A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 14. A1–A2. Thiarella bronni (Mayer, 1869), NHMW1864/0040/1123b, SL: 18.1 mm, MD: 6.6 mm, Borač (Boratsch) (the Czech Republic). B1–B2. Thiarella bronni (Mayer, 1869), NHMW1864/0040/1123a, Borač (Boratsch) (the Czech Republic). C1–C2. Thiarella bronni (Mayer, 1869), NHMW1855/0045/0045a, Hlohovec (the Czech Republic). D1–D2. Zafrona sphaerocorrugata nov. sp., NHMW 1866/0001/0814a, holotype, Niederleis (Austria). E. Zafrona sphaerocorrugata nov. sp., NHMW1866/0001/0814b, paratype, Niederleis (Austria). F1–F2. Zafrona sphaerocorrugata nov. sp., NHMW1866/0001/0814b, paratype, Niederleis (Austria). G1–G2. Scabrella haueri (Hoernes & Auinger, 1880), NHMW1863/0015/0632a, lectotype, Niederleis (Austria). H1–H2. Scabrella haueri (Hoernes & Auinger, 1880), NHMW1863/0015/0632b, paralectotype, Niederleis (Austria). I1–I2. Scabrella haueri (Hoernes & Auinger, 1880), NHMW1863/0015/0632, paralectotype, Niederleis (Austria).

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FIGURE 13A1–A2. OrthurellaconvexulaSacco,1904,NHMW1937 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 13A1–A2. OrthurellaconvexulaSacco,1904,NHMW1937/0002/0277,BadVöslau (Austria).B1–B2.Orthurellaconvexula Sacco, 1904, NHMW1846/0037/0271, Baden (Austria). C1–C2. Orthurella convexula Sacco, 1904, NHMW1856/0024/0013, Lăpugiu de Sus (Romania). D1–D2. Orthurella moravica (Hoernes & Auinger, 1880), NHMW1861/0011/0519a, paralectotype, Jaroměřice nad Rokytnou (the Czech Republic). E1–E2. Orthurella moravica (Hoernes & Auinger, 1880), NHMW1861/0011/ 0519b, lectotype, Jaroměřice nad Rokytnou (the Czech Republic). F1–F2. Orthurella moravica (Hoernes & Auinger, 1880), NHMW2020/0159/0001, paralectotype, Jaroměřice nad Rokytnou (the Czech Republic), G1–G2. Clinurella sophiae Boettger, 1902, SMF 360681, lectotype, Coşteiu de Sus (Romania). H1–H2. Orthurella convexula Sacco, 1904, NHMW1858/0043/0017, Lăpugiu de Sus (Romania). I. Orthurella moravica (Hoernes & Auinger, 1880), NHMW2020/0159/0002, paralectotype, Jaroměřice nad Rokytnou (the Czech Republic).

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FIGURE 9 A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 9 A1–A2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0001, holotype, Steinebrunn (Austria). B1–B2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0002, paratype, Steinebrunn (Austria). C1–C2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0003, paratype, Steinebrunn (Austria). D. Mitrella elongatissima nov. sp., NHMW1846/0037/0157, paratype, Steinebrunn (Austria). E1–E2. Mitrella demaintenonae nov. sp., NHMW1878/0001/0081, paratype, Möllersdorf; F1–F2. Mitrella demaintenonae nov. sp., NHMW2020/0162/0002, Coşteiu de Sus (Romania). G1–G2. Mitrella demaintenonae nov. sp., NHMW 2020/0161/0001, holotype, Coşteiu de Sus (Romania). H. Mitrella demaintenonae nov. sp., NHMW1867/ 0019/0049b, Coşteiu de Sus (Romania).

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FIGURE 12 A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 12 A1–A2. "Mitrella" bittneri (Hoernes & Auinger, 1880), NHMW1860/0001/0129a, lectotype, Steinebrunn (Austria). B1–B2. "Mitrella" bittneri (Hoernes & Auinger, 1880), NHMW1860/0001/0129b, paralectotype, Steinebrunn (Austria). C1– C2. "Mitrella" bittneri (Hoernes & Auinger, 1880), NHMW1860/0001/0129c, paralectotype, Steinebrunn (Austria). D1–D2. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1876/0011/0018, lectotype, Lăpugiu de Sus (Romania). E1–E2. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1859/0045/0601, paralectotype, Lăpugiu de Sus (Romania). F. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1854/0035/0133, paralectotype, Lăpugiu de Sus (Romania). G. Defensina bucciniformis (Hoernes & Auinger, 1880), SMF 360679, lectotype of Mitrella inquidens Boettger, 1906, Coşteiu de Sus (Romania).

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FIGURE 7 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 7. Species described by Friedberg (1911, 1938). A1–A2. "Columbella" paucicostata (Friedberg, 1911), ZNG PAN 1257, holotype, Dryszczów (Ukraine). B1–B2. Sulcomitrella januskiewiczi (Friedberg, 1938), ZNG PAN 2528, lectotype, Zborów (Ukraine). C2–C2. Mitrella convexa Friedberg, 1911, re-illustrated from Friedberg (1911, pl. 2, fig. 2).

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FIGURE 20 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 20. Schematic drawings of Paratethyan Columbellidae showing size relations amongst species; colors group genera. 1. Asytris? agenta Harzhauser & Kowalke, 2002, 2. Bellacolumbella katharinae (Hoernes & Auinger, 1880), 3. Bellacolumbella embryonalis (Boettger, 1902), 4. Mitrella pseudofallax (Kókay, 1966), 5. Bellacolumbella karreri (Hoernes & Auinger 1880), 6. Sulcomitrella januskiewiczi (Friedberg, 1938), 7. Costoanachis latecostata (Boettger, 1902), 8. Astyris sarmatica (Kojumdgieva, 1969), 9. Mitrella petersi (Hilber, 1879), 10. Costoanachis guembeli (Hoernes & Auinger, 1880), 11. Zafrona sphaerocorrugata nov. sp., 12. Defensina bucciniformis (Hoernes & Auinger, 1880), 13. Mitrella hilberi Cossmann, 1901, 14. "Mitrella" bittneri (Hoernes & Auinger, 1880, 15. Scabrella haueri (Hoernes & Auinger, 1880), 16. Mitrella demaintenonae nov. sp., 17. Mitrella elongatissima nov. sp., 18. Mitrella fallax (Hoernes & Auinger, 1880), 19. Auingeria austriaca (Hoernes & Auinger, 1880), 20. Clinurella sophiae Boettger, 1902, 21. Orthurella moravica (Hoernes & Auinger, 1880), 22. Thiarella bronni (Mayer, 1869), 23. "Columbella" paucicostata (Friedberg, 1911), 24. Mitrella dacica nov. sp., 25. Auingeria bellardii (Hoernes, 1852), 26. Auingeria grussbachensis (Csepreghy-Meznerics, 1969)., 27. Mitrella viennensis nov. sp., 28. Martaia zitteli (Hoernes & Auinger, 1880), 29. Mitrella convexa Friedberg, 1811, 30. Mitrella complanata (Sacco, 1890), 31. Graphicomassa korytnicensis (Bałuk, 1995), 32. Martaia dujardini (Hoernes & Auinger, 1880), 33. Columbella mayeri Hoernes, 1856, 34. Graphicomassa polonica (Pusch, 1837), 35. Orthurella convexula Sacco, 1904.

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FIGURE 6. A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 6. A1–A2. Costoanachis guembeli (Hoernes & Auinger, 1880), NHMW1887/0018/0021, Steinebrunn (Austria). B1–B2. Costoanachis latecostata (Boettger, 1902), SMF 360677, lectotype, Coşteiu de Sus (Romania). C1–C2. Costoanachis latecostata (Boettger, 1902), NHMW2020/0147/0001, Forchtenau (Austria). D1–D2. Costoanachis latecostata (Boettger, 1902), SMF 360679, lectotype of Columbella (Anachis) subcorrugata Boettegr, 1906, Coşteiu de Sus (Romania). E1–E2. Costoanachis latecostata (Boettger, 1902), NHMW1866/0001/1025a, Forchtenau (Austria).

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FIGURE 5. A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 5. A1–A2. Costoanachis guembeli (Hoernes & Auinger, 1880). NHMW1866/0001/1025b, lectotype, Steinebrunn (Austria). B1–B2. Costoanachis guembeli (Hoernes & Auinger, 1880), NHMW1949/0005/0007, paralectotype, Niederleis (Austria). C1–C2. Costoanachis guembeli (Hoernes & Auinger, 1880), NHMW1866/0001/1025c, paralectotype, Forchtenau (Austria). D1–D2. Costoanachis guembeli (Hoernes & Auinger, 1880), NHMW2020/0148/0001, Szob (Hungary). E1–E2. Costoanachis guembeli (Hoernes & Auinger, 1880), NHMW2020/0148/0002, Szob (Hungary).

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FIGURE 1. A in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

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FIGURE 16. A1–A2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 16. A1–A2. Martaia dujardini (Hoernes & Auinger, 1880), NHMW1858/0043/0018, lectotype, Lăpugiu de Sus (Romania). B1–B2. Martaia dujardini (Hoernes & Auinger, 1880), NHMW 1863/0015/0630a, paralectotype, Lăpugiu de Sus (Romania). C1–C2. Martaia dujardini (Hoernes & Auinger, 1880), NHMW1863/0015/0630b, paralectotype, SL: 17.9 mm, MD: 7.5 mm, Niederleis (Austria). D1–D2. Martaia zitteli (Hoernes & Auinger, 1880), NHMW1875/0009/0010b, paralectotype, Lăpugiu de Sus (Romania). E1–E2. Martaia zitteli (Hoernes & Auinger, 1880), NHMW1875/0009/0010a, paralectotype, Lăpugiu de Sus (Romania). F1–F2. Martaia zitteli (Hoernes & Auinger, 1880), NHMW1875/0009/0010c, lectotype, Lăpugiu de Sus (Romania).

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FIGURE 2 in The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea—striking diversity of a negelected group

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.

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FIGURE 3. A–B in New species of the genus Astyris (Gastropoda: Neogastropoda: Buccinoidea Columbellidae) from the vicinity of hydrothermal vents of the Piip Volcano southwestern Bering Sea

FIGURE 3. A–B. Radulae of studied Astyris spp. A. Astyris axicostata n. sp., specimen from the type locality. B. Astyris cf. rosacea(shell on Fig. 2 J). C–D. Astyris axicostata n. sp. in situ, specimens of Astyris axicostatus are marked with red arrows, insert shows enlarged fragment of the photos. C. Sta LV82-2/4, northern summit of Piip volcano, 55.24′57′′N, 167.16′35′′E, 384 м, 13.06.2018; rocks with bacterial mats. In addition to the Astyris, numerous isopods Eurycope andreyi (Munnopsidae) and Corallimorphus cf. pilatus (Corallimorpharia, Anthozoa) are seen. D. Type locality, rock with dense aggregations of Epizoanthus sp. (Zoantharia, Anthozoa).

opennotspecifiedApr 2023View details →
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FIGURE 2. Astyris axicostata n in New species of the genus Astyris (Gastropoda: Neogastropoda: Buccinoidea Columbellidae) from the vicinity of hydrothermal vents of the Piip Volcano southwestern Bering Sea

FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced.

opennotspecifiedApr 2023View details →
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FIGURE 1 in New species of the genus Astyris (Gastropoda: Neogastropoda: Buccinoidea Columbellidae) from the vicinity of hydrothermal vents of the Piip Volcano southwestern Bering Sea

FIGURE 1. Phylogenetic reconstruction using the COI gene (658 bp) showing Astyris axicostata n. sp. forming the clade with Astyris thermophila and A. rosacea nested within Columbellidae. Node values indicate Bayesian posterior probability and Maximal Likelihood bootstrap values, those lower than 0.8 and 80 are omitted.

opennotspecifiedApr 2023View details →
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FIG. 7 in On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species

FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm; E, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61639, SL 43.2 mm; F, AURORA 2007, st. CP2685, sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015, st. CP4133, not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm.

opencc-zeroFeb 2020View details →
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FIG. 3 in On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species

FIG. 3. — Phylogenetic tree of a dataset of Buccinoidea Rafinesque, 1815 obtained with Bayesian analysis of concatenated sequences of COI and 28S. Support values shown for the supported nodes with posterior probability values between 0.70 and 1.00 only.

opencc-zeroFeb 2020View details →
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FIG. 6 in On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species

FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm.

opencc-zeroFeb 2020View details →
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FIG. 9 in On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species

FIG. 9. — Shells of studied Siphonalia spp.: A-C, S. cassidariaeformis (Reeve, 1846): specimen no. 1 (A), specimen no. 2 (B), specimen no. 3 (C); D, E, S. pfefferi G. B. Sowerby III, 1900; D, specimen no. 1; E, specimen no. 2. Scale bar: 10 mm.

opencc-zeroFeb 2020View details →

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