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FIGURE 12 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 12. Pygidium and aedeagus (left view, dorsal view, right view) of species of Sauvagesinella, all scale bars 0.25 mm. A, Sauvagesinella becki; B, S. loftyensis new combination; C, S. monstrosa; D, S. palustris.

FIGURE 14 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 14. Habitus, prothoracic leg, and pygidium of male holotype of Lepanus ovatus (type species of Lepanus). A, habitus, scale 1 mm; B, protibia, scale 0.2 mm; C, pygidium, scale 0.2 mm.

FIGURE 11 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 11. Key characters of Sauvagesinella, scale bars 0.25 mm. A, lateral view of elytra (S. becki); B, ventral view of profemur, note flattened posterior face and the absence of trochantofemoral anterior pit (S. monstrosa); C, dorsal view of male protibia (S. becki ♂); D, ventral view of male protibia, arrow shows ornamentation (S. becki ♂); E, male metatarsus (S. monstrosa ♂).

FIGURE 10 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 10. Habitus of species of Monteithocanthon new genus in relative scale to each other, scale bars 1 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species.

FIGURE 8 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 8. Male metatibia of species of Monteithocanthon new genus, all scale bars 0.25 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species; G, female metathoracic leg of M. arator.

FIGURE 6 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 6. Habitus of species of Matthewsius new genus in relative scale to each other, scale bar 1 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species.

FIGURE 7 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba).

FIGURE 5 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 5. Distributional maps of species belonging to the new genera Matthewsius and Monteithocanthon including close up insets of A, Pilbara region, Western Australia; B, northeastern Queensland; C, southeastern Queensland; D, southwestern Western Australia; E, southeastern South Australia; F, central to southeastern New South Wales.

FIGURE 2 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 2. Key characters of Matthewsius new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. stricklandensis); B, ventral view of profemur, note presence of trochantofemoral anterior pit and ornamentation on apex of protibia (M. illawarrensis); C, close up of ornamentation on apex of protibia, white arrow pointing to comb and black arrow pointing to brush of setae (M. illawarrensis); D, prothoracic leg with three tibial teeth each with two groups of setae dorsally (M. illawarrensis ♂); E, male mesotibia with dense brush of setae (M. illawarrensis ♂); F, female mesotibia without dense brush of setae (M. illawarrensis ♀); G, male metatibia with small, blunt tooth (M. illawarrensis ♂); H, female metatibia without small, blunt tooth (M. illawarrensis ♀), inset includes close up of claw; I, male metatibia with small, blunt tooth (M. penelopae ♂).

FIGURE 4 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 4. Key characters of Aptenocanthon hopsoni, all scale bars 0.25 mm. A, lateral view of elytra, note presence of stria 8 remnant on pseudepipleuron; B, close up of basal angle of the elytra, note the presence of striae 8 and 9; C, ventral view of profemur, note presence of trochantofemoral anterior pit; D, pygidium; E, male mesotibia and tarsus; F, male metatibia and tarsus.

FIGURE 1 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 1. Phylogeny of Australian dung beetles. Topology generated from partitioned Bayesian analysis of Australasian dung beetles. Values at nodes represent posterior probability and bootstrap support for clades also supported in partitioned maximum likelihood analysis. Stars on nodes represent the position of the two new genera in comparison to Lepanus.

FIGURE 3 in Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species

FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species.

Fig. 9 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 9. Collection localities for Wadicosa benadira (Caporiacco) (stars), W. cognata sp. nov. (triangle) and W. jocquei sp. nov. (circles). One symbol may refer to more than one locality.

Fig. 6 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 6. Tegulum of left male palp with tegular apophysis (tg. a). Basal part and lower branch of tegular apophysis with spinulae (s) and verrucose outgrows (v), the latter enlarged in B and D. — A – B. Wadicosa cognata sp. nov., from Kenya, Lake Magadi. — C – D. W. Fdelis (Pickard-Cambridge), from Spain. Scale bars: A, C = 100 µm, B, D = 10 µm.

Fig. 3 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 3. Right male palp, ventral view (A – C) and epigyne (D – H). — A, D. Wadicosa benadira (Caporiacco) (A from Kenya, WNW of Mombasa; D from Kenya, Marsabit). — B, E. W. cognata sp. nov. B. Holotype. E. Paratype. — C, G. W. jocquei sp. nov. C. Holotype. G. Allotype. — F. W. Fdelis (Pickard-Cambridge) from Spain. — H. W. russellsmithi sp. nov., holotype. Scale bar: A – H = 0.5 mm.

Fig. 8 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 8. Epigyne in ventral (A – C) and dorsal (D – F) view. — A, D. Wadicosa benadira (Caporiacco), from Kenya, Marsabit. — B, E. W. cognata sp. nov., from Kenya, Lake Magadi. — C, F. W. jocquei sp. nov. C. From Mauritius. F. From Comoros, Anjouan. fo, foveola; l. el. lateral elevation; sp, spermatheca; arrow points at lateral elevation partly covering foveola. Scale bars: A – F = 300 µm.

Fig. 2. — A – D in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 2. — A – D. Wadicosa benadira (Caporiacco). A. Left male palp, from Caporiacco (1940: Fg. 1). B. Right male palp, from Roewer (1959: Fg. 18). C – D. Left embolus in Ƌ from Kenya, Marsabit; anterior (C) and ventral (D) view. — E – F. W. jocquei sp. nov. Left embolus (E), frontal view, and left cymbium (F) in Ƌ from Comoros, Anjouan. Arrow points at projecting retrolateral rim of cymbium. Scale bars: C – E = 0.2 mm, F = 0.5 mm.

Fig. 5 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 5. Terminal part of left male palp with embolus (em) and conductor (co). — A. Wadicosa cognata sp. nov., from Kenya, Lake Magadi, ventral view. — B. W. Fdelis (Pickard-Cambridge), from Spain, ventral view. — C – D. W. benadira (Caporiacco), from Kenya, WNW of Mombasa. C. Ventral view. D. More frontal view. — E. W. jocquei sp. nov., from Comoros, Moheli, frontal view. Scale bars: A – B = 200 µm, C – E = 200 µm.

Fig. 7 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 7. Epigyne, ventral view. — A. Wadicosa benadira (Caporiacco), from Kenya, Tsavo East NP. — B. W. cognata sp. nov., allotype. — C. W. jocquei sp. nov., from Madagascar, Mahafaly. — D. W. russellsmithi sp. nov., holotype. Scale bars = 0.2 mm.

Fig. 1 in Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa

Fig. 1. Habitus, dorsal view. — A – C. Wadicosa benadira (Caporiacco). A Ƌ from Kenya, Marsabit. B Ƌ and C ♀ from Kenya, WNW of Mombasa. — D. W. russellsmithi sp. nov., holotype, ♀. — E – F. W. cognata sp. nov. E Ƌ and F ♀ from Kenya, Lake Magadi. — G – H. W. jocquei sp. nov. G Ƌ from Comoros, Pomoni. H ♀ from Mayotte. Scale bar: A – H = 2 mm.