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Fig. 1. Phylogenetic hypothesis for Lyropaeus Waterhouse, 1878 in Molecular phylogeny and classification of Lyropaeini (Coleoptera: Lycidae) with description of larvae and new species of Lyropaeus
Fig. 1. Phylogenetic hypothesis for Lyropaeus Waterhouse, 1878 based on a maximum likelihood analysis of five fragments 18S and 28S rDNA, cox1, nad5 and rrnl mtDNA. Numbers at the branches are Bayesian frequencies and likelihood and parsimony bootstrap values.
Fig. 56 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; C–E, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo].
Fig. 47 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 47. Migmarma lustrum, new species, holotype male (15.7 × 13.9 mm) (ZRC 2010.0021), Irian Jaya, Indonesia. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of right chela; F, outer view of left chela; G, anterior thoracic sternum and pleon; H, pleon.
Fig. 42. A–E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 42. A–E, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo. A, male pleon; B, left G1 (ventral view, denuded); C, left G1 (dorsal view, denuded); D, left distal part of G1 (ventral view, denuded); E, left distal part of G1 (dorsal view, denuded). Scales: A = 2.0 mm; B, C = 1.0 mm; D, E = 0.5 mm.
Fig. 44. Vulva. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 44. Vulva. A, Pseudosesarma edwardsii, female (16.5 × 15.3 mm) (ZRC 2016.0608), Malaysia; B, Pseudosesarma crassimanum, female (15.9 × 13.8 mm) (ZRC 2000.1768), Malaysia; C, Pseudosesarma crassimanum, female (18.0 × 15.7 mm) (ZRC 2017.1045), Thailand; D, Pseudosesarma anteactum, paratype female (17.5 × 15.4 mm) (ZRC 2016.0603), Sri Lanka; E, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, Pseudosesarma boulengeri, female (24.2 × 20.0 mm) (ZRC 2014.0335), Iran; G, Contusarma bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Thailand; H, Contusarma cheirogonum, female (26.6 × 23.0 mm) (ZRC 1964.9.28.127), Sarawak; I, Miersarma granosimanum, female (20.1 × 16.4 mm) (ZRC 1965.7.29.166), Malaysia; J, Manarma moeschii, female (19.3 × 17.3 mm) (ZRC 2017.0170), Thailand.
Fig. 53 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 53. Colour in life. A, Orisarma dehaani, male (23.4 × 20.8 mm) (ZRC 2012.0057), Kagoshima, Japan [photograph: PKL Ng]; B, O. dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; C, O. dehaani, male (27.6 × 29.6 mm) (ZRC 2002.0223), Hong Kong [photograph: Tan Heok Hui]; D, O. dehaani, male (26.5 × 24.0 mm) (ZRC 2019.0539), Hong Kong [photograph: Paul Ng]; E, O. dehaani, female (20.2 ×17.6 mm) (ZRC 2019.0539), Hong Kong [photograph: Paul Ng]; F, O. neglectum, Shanghai, China, specimens not preserved [photograph: Liu Wenliang].
Fig. 57 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 57. Colour in life. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand [photograph: Phaibul Naiyanetr]; B–D, C. cheirogonum, male (26.3 × 23.0 mm) (ZRC 1995.0226), Bako, Sarawak [photograph: PKL Ng]; E, G, C. cheirogonum, male, Tioman, Peninsular Malaysia, specimen not collected [photograph: Chua Ee Kiam]; F, C. cheirogonum, Tioman, Peninsular Malaysia, female (specimen not collected); H, C. cheirogonum, specimen not collected, Bako, Sarawak [photograph: Jongkar Grinang].
Fig. 38 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 38. Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand. A, anterior thoracic sternites 1–4; B, left third maxilliped (denuded); C, male pleon; D, left G1 (ventral view, denuded); E, left G1 (dorsa view); F, left distal part of G1 (ventral view, denuded); G, left distal part of G1 (dorsal view, denuded); H, left distal part of G1 (mesial view). Scales: A, C = 5.0 mm; B = 2.5 mm; D, E = 1.0 mm; F–H = 0.5 mm.
Fig. 41. A–E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 41. A–E, Manarma moeschii, male (20.2 × 18.0 mm) (ZRC 2000.1926), Thailand; F–J, M. johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore. A, F, male pleon; B, G, left G1 (ventral view, denuded); C, H, left G1 (dorsal view, denuded); D, I, left distal part of G1 (ventral view, denuded); E, J, left distal part of G1 (dorsal view, denuded). Scales: A = 5.0 mm; B, C, G, H = 1.0 mm; F = 2.0 mm; D, E, I, J = 0.5 mm.
Fig. 35. A–E, Pseudosesarma crassimanum. A–G in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 35. A–E, Pseudosesarma crassimanum. A–G, lectotype male (16.3 × 14.6 mm) (NHM 1886.52b), Myanmar; H–M, male (15.9 × 14.3 mm) (ZRC 2008.0442), Ranong; N–R, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore. A, N, pleon; B–F, right G1; G, right G2; H–L, O–R, left G1; M, left G2. Scales: A = 2.0 mm; B, G, H, I, M, O, P = 1.0 mm; N = 5.0 mm; C–F, J–L, Q, R = 0.5 mm. After Ng & Schubart (2017: fig. 11).
Fig. 40. Miersarma granosimanum. A–E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 40. Miersarma granosimanum. A–E, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; F–I, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; J, male (19.5 × 16.7 mm) (ZRC 1965.7.29.165), Sedili River, Johor, Malaysia. A, male pleon; B, F, left G1 (ventral view, denuded); C, G, J, left G1 (dorsal view, denuded); D, H, left distal part of G1 (ventral view, denuded); E, I, left distal part of G1 (dorsal view, denuded). Scales: A = 5.0 mm; B, C, F, G, J = 1.0 mm; D, E, H, I = 0.5 mm.
Fig. 33. A–E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 33. A–E, male anterior thoracic sternum and pleon; F–H, left ambulatory leg. A, D, H, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; B, E, M. johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; C, F, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; G, O. patshuni (, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong.
Fig. 32. Male sternopleonal cavity. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 32. Male sternopleonal cavity. A, Pseudosesarma edwardsii, male (20.8 × 18.6mm (ZRC 2003.0084), Pulau Tekong, Singapore; B, Pseudosesarma crassimanum, male (15.1 × 13.3 mm) (ZRC 2000.1768), Sungei Benut, Johor, Malaysia; C, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; D, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia. Arrow indicates shallow depression on somite 5 to accept G1 tip when pleon closed (depression absent in Miersarma, D).
Fig. 46 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 46. Migmarma modestum, holotype male (25.0 × 20.9 mm) (SMF 1989), Indonesia. A, left third maxilliped (denuded); B, pleon; C, left G1 (ventral view); D, left G1 (dorsal view); E, distal part of left G1 (ventral view); F, distal part of left G1 (mesial view); G, distal part of left G1 (dorsal view); H, distal part of left G1 (mesial view); I, left G2. Scales: A, C, D, I = 1.0 mm; B = 5.0 mm; E–H = 0.5 mm.
Fig. 55 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 55. Colour in life. A, Orisarma patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong [photograph: PKL Ng]; B–D, O. patshuni, male (13.8 × 12.3 mm) (ZRC 2019.1067), Hong Kong [photograph: Paul Ng].
Fig. 49 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 49. Migmarma lustrum, new species, holotype male (15.7 × 13.9 mm) (ZRC 2010.0021), Irian Jaya, Indonesia. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, distal part of left G1 (mesial view); F, left G2. Scales = 0.5 mm.
Fig. 34. Pseudosesarma edwardsii. A, B, D, E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 34. Pseudosesarma edwardsii. A, B, D, E, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; C, male (13.3 × 12.9 mm) (ZRC 2000.2019), Pulau Ubin, Singapore; F–H, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi; I, J, male (21.3 × 18.7 mm) (ZRC 2003.0084), Singapore; K, L, male (17.8 × 16.3 mm) (ZRC 2016.0608), Langkawi; M, N, male (12.5 × 10.4 mm) (ZRC 2016.0608), Langkawi. A, anterior thoracic sternites 1–4; B, C, male pleon; D, H, I, K, M, left G1 (dorsal view, denuded); E, G, J, K, N, left G1 (ventral view, denuded); H, left G2. Scales: A–C = 2.0 mm; D–J = 1.0 mm; K–N = 0.5 mm. (After Ng & Schubart, 2017: figs. 6, 7).
Fig. 43. Vulva. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 43. Vulva. A, Chiromantes haematocheir, female (27.0 × 23.7 mm) (ZRC 2002.0225), Japan; B, Orisarma dehaani, female (20.4 × 17.8 mm) (ZRC 2012.0057), Japan; C, Orisarma intermedium, female (31.8 × 28.4 mm) (ZRC 2013.0140), Japan; D, Orisarma patshuni, female (20.5 × 18.1 mm) (ZRC 2014.0817), Hong Kong; E, Danarma obtusifrons, female (15.7 × 11.6 mm) (UF-FLMNH 14837), Hawaii; F, Danarma eurymerus, paratype ovigerous female (18.0 × 13.4 mm) (ZRC 2012.0956), Taiwan; G, Cristarma eulimene, female (15.6 × 12.8 mm) (ZRC 1968.1.22.3), Mozambique; H, Cristarma ortmanni, female (13.1 × 10.2 mm) (ZRC 2000.1783), Kenya; I, Trapezarma angolense, female (18.2 × 14.3 mm) (ZRC 2015.0297), Cameroon; J, Platychirarma buettikoferi, ovigerous female (11.3 × 9.1 mm) (ZRC 2015.0298), Cameroon.
Fig. 31. Male anterior thoracic sternum and pleon. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 31. Male anterior thoracic sternum and pleon. A, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; B, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; C, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; D, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; E, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; F, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; G, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; H, Bresedium laevimanum, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia.
Fig. 29 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species
Fig. 29. Outer view of chela. A, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; B, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; C, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; D, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; E, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; F, Bresedium laevimanum, male (18.1 × 16.4 mm) (ZRC 1972.3.7.25), Sarawak.
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