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Fig. 7 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 7. Parameres and median lobe in ventral view. a. Strigoderma villosella (Blanchard, 1851). b–f. Strigoderma spp. Scale bars = 0.5 mm.

Fig. 3 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 3. Tegmen with the parameres in perpendicular position. a. Strigoderma villosella (Blanchard, 1851). b. Moroniella Ramírez-Ponce, 2015. c. Balanogonia Paucar-Cabrera, 2003. d. Callirhinus Blanchard, 1851. e. Mazahuapertha Morón & Nogueira, 1998. Scale bars = 0.5 mm.

Fig. 2 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 2. Dorsal habitus of American genera of Anomalini Peringuey, 1902 with their corresponding tegmen in ventral view. Scale bars: specimen = 2 mm; tegmen = 0.5 mm.

Fig. 6 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 6. Dorsal habitus of the genera with perpendicular parameres. a. Strigoderma villosella (Blanchard, 1851). b. Moroniella Ramírez-Ponce, 2015. c. Balanogonia Paucar-Cabrera, 2003. d. Callirhinus Blanchard, 1851. e. Mazahuapertha Morón & Nogueira, 1998. Scale bars = 1 mm.

Fig. 1. Aedeagus and its parts. a–b in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 1. Aedeagus and its parts. a–b. Paranomala corcovada Ramírez-Ponce et al., 2014. a. Lateral view. b. Dorsal view. c–l. Tegmen in dorsal and lateral view. c–d. Paranomala inbio Ramírez-Ponce et al., 2014. e–f. Paranomala strigodermoides (Filippini et al., 2015). g–h. Epectinaspis moreletiana (Blanchard, 1851). i–j. Strigoderma ngabe Ramírez-Ponce & Curoe 2014. k–l. Paranomala Casey, 1915. Scale bar = 0.5 mm. Abbreviations: BP = basal piece; IS = internal sac; ISarm = internal sac armature; ML = median lobe; PM = parameres.

Fig. 9 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 9. Distribution map of the genera with perpendicular parameres (the only genus not endemic to Mexico is Moroniella Ramírez-Ponce, 2015, which extends into Central America).

Fig. 8 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 8. Morphological characters in Lamoana villosella (Blanchard, 1851). a. Ventral view of habitus. b. Oblique view of habitus.c. Metasternum.d. Dorsal view of head and pronotum. e. Protibia.f. Protarsus.

Fig. 4 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 4. Parameres (caudal view) and median lobe (ventral view). a–b. Strigoderma villosella (Blanchard, 1851). c–d. Moroniella Ramírez-Ponce, 2015. e–f. Balanogonia Paucar-Cabrera, 2003. g–h. Callirhinus Blanchard, 1851. i–j. Mazahuapertha Morón & Nogueira, 1998. Scale bars = 0.5 mm.

Fig. 5 in Comparative morphology of the genera with perpendicular parameres in Anomalini (Coleoptera, Scarabaeidae, Rutelinae) allows revalidation of the endemic Mexican genus Lamoana Casey, 1915 stat. rev.

Fig. 5. Lateral view of the genera with perpendicular parameres. a. Strigoderma villosella (Blanchard, 1851). b. Moroniella Ramírez-Ponce, 2015. c. Balanogonia Paucar-Cabrera, 2003. d. Callirhinus Blanchard, 1851. e. Mazahuapertha Morón & Nogueira, 1998. Scale bars = 1 mm.

Рис. 23–31. Tonkinilla graminea, gen. et comb. nov., синтип: 23–25, общий виΔ; 26 — верхняя губа, наΛичник и чеΛюстные щупики; 27 — наΔкрыΛья сбоку; 28, 29 — простернаΛьный отросток (28 — сбоку; 29 — спереΔи); 30, 31 — парамеры и базаΛьная пΛастинка эΔеагуса (30 — виΔ сверху; 31 — виΔ снизу). Масштаб: 23–25 — 2,5 мм; 26, 27, 30, 31 — 1 мм; 28 — 0,5 мм; 29 — 0,3 мм (Λинейка общая ΔΛя 23–25 и ΔΛя 30 и 31) Figs. 23–31. Tonkinilla graminea, gen. et comb. nov., syntype: 23–25, habitus; 26 — labrum, clypeus and mandibular palpi; 27 — lateral view of elytron; 28, 29 — prosternal process (28 — lateral view; 29 — frontal view); 30, 31 — parameres and basal plate of aedeagus (30 — dorsal view; 31 — ventral view). Scale bars: 23–25 — 2.5 mm; 26, 27, 30, 31 — 1 mm; 28 — 0.5 mm; 29 —0.3 mm (common bar for 23–25 and for 30 and 31) in On the systematics of the Anomalina subtribe in Southeast Asia (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

Рис. 23–31. Tonkinilla graminea, gen. et comb. nov., синтип: 23–25, общий виΔ; 26 — верхняя губа, наΛичник и чеΛюстные щупики; 27 — наΔкрыΛья сбоку; 28, 29 — простернаΛьный отросток (28 — сбоку; 29 — спереΔи); 30, 31 — парамеры и базаΛьная пΛастинка эΔеагуса (30 — виΔ сверху; 31 — виΔ снизу). Масштаб: 23–25 — 2,5 мм; 26, 27, 30, 31 — 1 мм; 28 — 0,5 мм; 29 — 0,3 мм (Λинейка общая ΔΛя 23–25 и ΔΛя 30 и 31) Figs. 23–31. Tonkinilla graminea, gen. et comb. nov., syntype: 23–25, habitus; 26 — labrum, clypeus and mandibular palpi; 27 — lateral view of elytron; 28, 29 — prosternal process (28 — lateral view; 29 — frontal view); 30, 31 — parameres and basal plate of aedeagus (30 — dorsal view; 31 — ventral view). Scale bars: 23–25 — 2.5 mm; 26, 27, 30, 31 — 1 mm; 28 — 0.5 mm; 29 —0.3 mm (common bar for 23–25 and for 30 and 31)

Рис. 16–22. Anomala malivirens sp. nov., гоΛотип (16–19), A. cribrata, Ява (cZ) (20), A. diana, Ассам (cZ) (21) и A. perplexa, синтип, Сикким (NHM) (22): 16 — общий виΔ; 17, 20–22 — эΔеагус сбоку; 18 — парамеры сверху; 19 — парамеры и базаΛьная пΛастика снизу. Масштаб: 16 — 5 мм; 17–22 — 1 мм (Λинейка общая ΔΛя 18 и 19) Figs. 16–22. Anomala malivirens sp. nov., holotype (16–19), A. cribrata, Java (cZ) (20), A. diana, Assam (cZ) (21) и A. perplexa, syntype, Sikkim (NHM) (22): 16 — habitus; 17, 20–22 — aedeagus, lateral view; 18 — parameres, dorsal view; 19 — parameres and basal plate, ventral view. Scale bars: 16 — 5 mm; 17–22 — 1 mm (common bar for 18 and 19) in On the systematics of the Anomalina subtribe in Southeast Asia (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

Рис. 16–22. Anomala malivirens sp. nov., гоΛотип (16–19), A. cribrata, Ява (cZ) (20), A. diana, Ассам (cZ) (21) и A. perplexa, синтип, Сикким (NHM) (22): 16 — общий виΔ; 17, 20–22 — эΔеагус сбоку; 18 — парамеры сверху; 19 — парамеры и базаΛьная пΛастика снизу. Масштаб: 16 — 5 мм; 17–22 — 1 мм (Λинейка общая ΔΛя 18 и 19) Figs. 16–22. Anomala malivirens sp. nov., holotype (16–19), A. cribrata, Java (cZ) (20), A. diana, Assam (cZ) (21) и A. perplexa, syntype, Sikkim (NHM) (22): 16 — habitus; 17, 20–22 — aedeagus, lateral view; 18 — parameres, dorsal view; 19 — parameres and basal plate, ventral view. Scale bars: 16 — 5 mm; 17–22 — 1 mm (common bar for 18 and 19)

Рис. 6–15. Anomala macrorhynchos sp. nov., гоΛотип (6–9), A. millestriga, синтип (?), «Sze-chuen» (ZMB) (10–12) и A. punctissima, паратип, «Laos, Vientiane prov., Ban Van Eue» (cF) (13–15): 6 — общий виΔ; 7, 10, 13 — эΔеагус, виΔ сбоку; 8, 11, 14 — парамеры, виΔ сверху; 9, 12, 15 — парамеры и базаΛьная пΛастинка, виΔ снизу. Масштаб: 6 — 4 мм; 7–15 — 1 мм (Λинейка общая ΔΛя 8 и 9, ΔΛя 11 и 12 и ΔΛя 14 и 15) Figs. 6–15. Anomala macrorhynchos sp. nov., holotype (6–9), A. millestriga, syntype (?), "Sze-chuen" (ZMB) (10–12), and A. punctissima, paratype, "Laos, Vientiane prov., Ban Van Eue" (cF) (13–15): 6 — habitus; 7, 10, 13 — aedeagus, lateral view; 8, 11, 14 — parameres, dorsal view; 9, 12, 15 — parameres and basal plate, ventral view. Scale bars: 6 — 4 mm; 7–15 — 1 mm (common bar for 8 and 9, for 11 and 12 and for 14 and 15) in On the systematics of the Anomalina subtribe in Southeast Asia (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

Рис. 6–15. Anomala macrorhynchos sp. nov., гоΛотип (6–9), A. millestriga, синтип (?), «Sze-chuen» (ZMB) (10–12) и A. punctissima, паратип, «Laos, Vientiane prov., Ban Van Eue» (cF) (13–15): 6 — общий виΔ; 7, 10, 13 — эΔеагус, виΔ сбоку; 8, 11, 14 — парамеры, виΔ сверху; 9, 12, 15 — парамеры и базаΛьная пΛастинка, виΔ снизу. Масштаб: 6 — 4 мм; 7–15 — 1 мм (Λинейка общая ΔΛя 8 и 9, ΔΛя 11 и 12 и ΔΛя 14 и 15) Figs. 6–15. Anomala macrorhynchos sp. nov., holotype (6–9), A. millestriga, syntype (?), "Sze-chuen" (ZMB) (10–12), and A. punctissima, paratype, "Laos, Vientiane prov., Ban Van Eue" (cF) (13–15): 6 — habitus; 7, 10, 13 — aedeagus, lateral view; 8, 11, 14 — parameres, dorsal view; 9, 12, 15 — parameres and basal plate, ventral view. Scale bars: 6 — 4 mm; 7–15 — 1 mm (common bar for 8 and 9, for 11 and 12 and for 14 and 15)

Рис. 1–5. Anomala hamuliphalla sp. nov.: 1, 2 — общий виΔ (1 — гоΛотип, самец; 2 — паратип, самка); 3–5 — эΔеагус (3 — сбоку; 4 — парамеры сверху; 5 — парамеры и базаΛьная пΛастинка снизу). Масштаб: 1, 2 — 2,5 мм (Λинейка общая); 3 — 1,0 мм; 4, 5 — 0,5 мм (Λинейка общая) Figs. 1–5. Anomala hamuliphalla sp. nov.: 1, 2 — habitus (1 — holotype, male; 2 — paratype, female); 3–5 — aedeagus (3 — lateral view; 4 — parameres, dorsal view; 5 — parameres and basal plate, ventral view). Scale bars: 1, 2 — 2.5 mm (common bar); 3 — 1.0 mm; 4, 5 — 0.5 mm (common bar) in On the systematics of the Anomalina subtribe in Southeast Asia (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

Рис. 1–5. Anomala hamuliphalla sp. nov.: 1, 2 — общий виΔ (1 — гоΛотип, самец; 2 — паратип, самка); 3–5 — эΔеагус (3 — сбоку; 4 — парамеры сверху; 5 — парамеры и базаΛьная пΛастинка снизу). Масштаб: 1, 2 — 2,5 мм (Λинейка общая); 3 — 1,0 мм; 4, 5 — 0,5 мм (Λинейка общая) Figs. 1–5. Anomala hamuliphalla sp. nov.: 1, 2 — habitus (1 — holotype, male; 2 — paratype, female); 3–5 — aedeagus (3 — lateral view; 4 — parameres, dorsal view; 5 — parameres and basal plate, ventral view). Scale bars: 1, 2 — 2.5 mm (common bar); 3 — 1.0 mm; 4, 5 — 0.5 mm (common bar)

FIGURES 48–53 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 48–53. Lateral view of Pachystethus species: 48) P. viduus a) male, b) female; 49) P. nectoctenus a) male, b) female; 50) P. crassesculptus male; 51) P. n u t a n s male; 52) P. matzapatlecus male; 53) P. ixtacomitanus male. Scale lines 1 mm.

FIGURES 42–47 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 42–47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm.

FIGURES 36–41 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 36–41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm.

FIGURES 2–7 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 2–7. Diagnostic structures in P. viduus: 2) right proleg; 3) right antenna, 4) male genitalia in dorsal view, 5) male genitalia in lateral view, 6) spiculum gastrale, 7) female genital plates. Scale lines 1 mm.

FIGURES 8–13 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 8–13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm.

FIGURES 20–25 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 20–25. Diagnostic structures in P. nectoctenus: 20) right proleg, 21) right antenna, 22) male genitalia in dorsal view, 23) male genitalia in lateral view, 24) spiculum gastrale, 25) female genital plates. Scale lines 1 mm.

FIGURES 14–19 in Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

FIGURES 14–19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm.