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Figure 10. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 10. Ebertichthys ettlingensis n. gen. et n. sp. Pelvic plate and pelvic fin (JME ETT 108). Abbreviations: Ant, anterior; pel.r, pelvic rays; p.pl, pelvic plate or basipterygium; p.sp, pelvic splint.

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Figure 3. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 3. Ebertichthys ettlingensis n. gen. et n. sp. in lateral view (holotype JME ETT 108). (a) Photograph of cranium and anterior part of body. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Drawing of cranium and anterior part of body. Abbreviations: ant, antorbital; br, branchiostegal rays; cl, cleithrum; cor, coracoid; dpa, dermopalatine; ent, entopterygoid; exc, extrascapula; hy, hyomandibula; io1-5, infraorbitals 1–5; iop, interopercle; l.de, left dentary; leth, lateral ethmoid; l.pmx, left premaxilla; met, mesethmoid; mx, maxilla; op, opercle; orb, orbitosphenoid; pa[= fr], parietal [= frontal bone of traditional terminology]; par, parasphenoid; pcl1-3, postcleithra 1–3; pop, preopercle; ppa[= pa], postparietal [= parietal bone of traditional terminology]; pt, pterotic; ptsp, pterosphenoid; ptt, posttemporal; qu, quadrate; sc, scales; sca, scapula; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sn, supraneurals; sorb.b, broken supraorbital bone; sop, subopercle; sy, symplectic; r.de, right dentary; r.pmx, right premaxilla; vo, vomer.

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Figure 19 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 19. Ascalabos voithii. Caudal skeleton in lateral view with only two epaxial basal fulcra preserved (neotype, JME SOS 537). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; dpr, dorsal or epaxial procurrent rays; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–4, hypurals 1–4; hbfu, hypaxial basal fulcra; hsPU4, haemal spine of preural centrum 4; naPU1, neural arch of preural centrum 1; nsPU2–4, neural spine of preural centra 2–4; PH, parhypural; PU1,4, preural centrum 1, 4; rer, rudimentary fin ray; sc, dorsal and ventral caudal scutes; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; 'UD', "urodermals"; UN1–8, uroneurals 1–8; PR1–19, principal rays 1–19; vpr, ventral procurrent rays; v.scu, ventral caudal scute.

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Figure 11. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 11. Ebertichthys ettlingensis n. gen. et n. sp. (JME ETT 64a). (a) Details of first two dorsal pterygiophores and first dorsal-fin rays. (b) Detail of last dorsal pterygiophores. Arrows point to the two articular areas for fin rays. Abbreviations: Ant, anterior; 1st d.pt, first dorsal pterygiophore; 2nd d.pt, second dorsal pterygiophore; ld.pt, last dorsal pterygiophore.

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Figure 4. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 4. Ebertichthys ettlingensis n. gen. et n. sp. Crania in lateral views illustrating specific morphological features. (a) JME ETT 60. (b) JME ETT 24. Arrow points to a small toothed patch. Abbreviations: ang, angular; aup, autopalatine; b.ant, broken antorbital; a.cer, anterior ceratohyal; br, branchiostegal rays; cl, cleithrum; de, dentary; ect, ectopterygoid; ent, entopterygoid; ghy, glossohyal; d.hy, dorsal hypohyal; iop, interopercle; hy, hyomandibula; io1–5, infraorbitals 1–5; l.ang, left angular; l.de, left dentary; l.pmx, left premaxilla; met, mesethmoid; mtg, metapterygoid; mx, maxilla; op, opercle; par, parasphenoid; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; ra, remnant of a branchial arch; r.de, right dentary; r.pmx, right premaxilla; scl, supracleithrum; scl.b, broken sclerotic bones; smx1–2, supramaxillae 1–2; sorb, supraorbital; sop, subopercle; ur, urohyal; v.hy, ventral hypohyal; vo, vomer.

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Figure 18 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 18. Ascalabos voithii. Caudal skeleton in lateral view slightly modified from Arratia (1997). Abbreviations: Ant, anterior; dpr, dorsal or epaxial procurrent rays; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; E1–3, epurals 1–3; H1–11, hypurals 1–11; hsPU2, haemal spine of preural centrum 2; mo, membranous outgrowth on anterodorsal margin of first uroneural; mo', membranous outgrowth on anterodorsal margin of second uroneural; nsPU2–4, neural spine of preural centra 2–4; PR1,19, principal caudal rays 1, 19; rer, rudimentary fin ray; U1+2+H1–2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; 'UD', "urodermals"; UN1–7, uroneurals 1–7; v.scu., ventral caudal scute.

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Figure 9. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 9. Ebertichthys ettlingensis n. gen. et n. sp. (a) Photograph of young specimen (JME ETT 61) in lateral view. Photograph courtesy of H. Tischlinger, Scale equals 1 cm. (b) Detail of last caudal vertebrae illustrating pseudo-diplospondyly and caudal endoskeleton incompletely preserved. Abbreviations: chcPU4, chordacentrum of preural centrum 4; chcU1–3, chordacentra of ural centra 1–3; E, epurals (the first one is missing); H1–7, hypurals 1–7; hsPU2, haemal spine of preural centrum 2; PH, parhypural; 'UD', "urodermal"; UN1–6, uroneurals 1–6.

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Figure 2. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 2. Ebertichthys ettlingensis n. gen. et n. sp. in lateral views. (a) Holotype JME ETT 108. (b) Paratype JME ETT 132a. (c) Paratype JME ETT 24, specimen with a fish prey in its mouth (small individual of Orthogonikleithrus). Scales equal 1 cm. Photographs courtesy of M. Ebert.

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Figure 13. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 13. Ebertichthys ettlingensis n. gen. et n. sp. Caudal fin and endoskeleton in lateral view (JME ETT 60). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; ffu, fringing fulcra; H1–6, hypurals 1–6; hsPU4, haemal spine of preural centrum 4; mo, membranous outgrowth on anterodorsal margin of first uroneural; nsPU1–4, neural spine of preural centra 1–4; PU1,4, preural centrum 1, 4; U1+2, ural centrum 1 + 2 (polyural terminology) fused with the bases of hypurals 1 and 2; 'UD', "urodermal"; UN1–5, uroneurals 1–5; PR1–19, principal rays 1–19; v.csu, ventral caudal scute.

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Figure 1 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 1. Distribution of Plattenkalk basins and reef areas in the southern Franconian Alb during the Early Tithonian (slightly modified from Viohl, 1996). The new fish described herein was recovered in Ettling.

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Figure 17 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 17. Ascalabos voithii (neotype, JME SOS 537). Midcaudal vertebrae illustrating the presence of a lateral crest and grooves on the lateral side of the vertebral centra. Arrows point to a lateral longitudinal crest. Abbreviations: na, neural arch; ns, neural spine; ha, haemal arch; hs, haemal spine; vc, vertebral centrum.

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Figure 14 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 14. Ascalabos voithii (neotype, JME SOS 537). (a) Photograph of neotype in lateral view. Photograph courtesy of M. Ebert. Scale equals 1 cm. (b) Cranium in lateral view and enlargement of premaxilla (c). Abbreviations: ang, angular; cl, cleithrum; de, dentary; ent, entopterygoid; io1–3, infraorbitals 1–3; mx, maxilla; par, parasphenoid; pcl1, postcleithrum 1; pmx, premaxilla; pop, preopercle; ptt, posttemporal; qu, quadrate; scl, supracleithrum; scl.b, broken sclerotic bone; smx1–2, supramaxillae 1–2; sop, subopercle; sy, symplectic.

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Figure 12. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 12. Ebertichthys ettlingensis n. gen. et n. sp. Caudal fin and endoskeleton in lateral view (holotype JME ETT 108). Abbreviations: Ant, anterior; d.scu, dorsal caudal scute; ebfu, epaxial basal fulcra; ffu, fringing fulcra; H1–7, hypurals 1–7; hsPU4, haemal spine of preural centrum 4; nsPU2–4, neural spines of preural centra 2–4; naU1+2, neural arches of ural centrum 1 + 2; PH, parhypural; prp, procurrent rays; PU1, 4, preural centra 1, 4; U1+2, 3, ural centrum 1 + 2 and 3 (polyural terminology) fused with the bases of hypurals 1 and 2; 'UD', "urodermal"; UN1–4, uroneurals 1–4; PR1–19, principal rays 1–19; v.csu, ventral caudal scute.

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Figure 7. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 7. Ebertichthys ettlingensis n. gen. et n. sp. Dorsal and anal fins and their relationships to the vertebral column (JME ETT 132a). Abbreviations: Ant, anterior; an.r, principal anal rays; epl.b, epipleural bones; dor.r, dorsal principal rays; epin.p, epineural processes; ha, haemal arch; hs, haemal spine; la.pt, last anal pterygiophore; ld.pt, last dorsal pterygiophore; na, neural arch; ns, neural spine; paph, parapophyses; pr.anr, anal procurrent rays; pr.dr, dorsal procurrent rays; ri, ribs; sn, supraneural bones; V21, vertebra 21; 1st a.pt, first anal pterygiophore; 1st d.pt, first dorsal pterygiophore.

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Figure 6. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 6. Ebertichthys ettlingensis n. gen. et n. sp. Details of certain abdominal vertebrae and pectoral fin. (a) Anterior abdominal region and pectoral girdle and fin (holotype JME ETT 108). (b) Enlargement of a few abdominal vertebrae illustrating details of the epineural processes (holotype JME ETT 108). (c) Details of pectoral fin and radials (JME ETT 132a). Abbreviations: cl, cleithrum; cor, coracoid; d.ra, distal radials; epin.p, epineural process; na, neural arch; ns, neural spine; paph, parapophyses; pcl1–3, postcleithrum 1–3; pec.f, pectoral fin; pec.ra, proximal radials supporting pectoral rays; pel.p, pelvic plate; ptt, posttemporal; ri, ribs; sc, anterior field of cycloid scales; sca, scapula; scl, supracleithrum; sn, supraneurals; vc, abdominal vertebral centra; 1st. pec, first pectoral ray.

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Figure 15 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 15. Ascalabos voithii from Schernfeld in the Eichstätt Basin. Uncatalogued specimen from U. Eller (Dümpelfeld). (a) Photograph of specimen in lateral view. (b) Enlargement of the head. Note the peculiar shape and distribution of the sensory tubules of the preopercular canal. Photographs courtesy of U. Eller.

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Figure 5. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 5. Ebertichthys ettlingensis n. gen. et n. sp. Infraorbital bones, preopercle, suspensorium, and part of upper jaw (JME ETT 148). Abbreviations: hy, hyomandibula; io1–4, infraorbitals 1–4; io5?, broken infraorbital 5?; mtg, metapterygoid; mx, maxilla; pop, preopercle; qu, quadrate; smx1–2, supramaxillae 1–2.

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Figure 20 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 20. Hypothesis of phylogenetic relationships of the Late Jurassic Ascalabos and Ebertichthys n. gen. among the most primitive teleosts (highlighted in bold). Synapomorphies supporting the main nodes are listed below; for a complete list see Arratia (2013, 114– 118). Uniquely derived characters are identified with an asterisk [*]. See Supplement 1 for descriptions of characters listed below. Node A (teleosteomorphs or total-group teleost): 10[0]*, 24[1]*, 40[0]*, 47[2], 50[1], 94[0]*, 108[1], 112[1], 142[1]*, 164[1]*, 165[1]*, 168[1]*, and 169[1]*. Characters 142 and 164–168 are soft anatomical features that are unknown in fossils, but the parsimony analysis predicts that they were present at this phylogenetic level. Node B: 25[1], 34[1], 53[1], 55[1], 56[1], 59[1], 100[1], 109[1], 122[1], 125[1], and 162[1]. Node C: 47[1], 48[1], 59[2], 62[1], 63[1], 65[1], 69[1], 118[1], 148[2], and 153[1]. Node D: 78[1]*, 79[1], 102[1], 132[1], 150[1], and 151[1]. Node E: 13[1], 18[1], 56[0], 80[1]*, 96[1], 110[1], and 131[1]. Node F: 48[0], 66[1], 70[1], 82[1]*, 105[1], 108[0], 109[0], 127[1], and 129[1]. Node G: 3[1], 25[0], 42[1], 64[1], 68[1]*, 108[3], 111[1]*, and 162[0]. Node H: 28[1], 30[1], 32[1], 56[1], 67[1], 69[2], 83[1]*, 100[0], 103[1]*, 121[1], 124[1], 140[1], and 149[1]*. Character 103[1] was incorrectly listed as a synapomorphy of node E in Arratia (2013). Node I: 50[2], 107[1], 114[1], 119[1], 120[1], 144[1], 148[3], 152[1], 155[1], and 156[2]. Node J (Ascalaboidae, n. fam.): 25[1], 29[1]*, 30[2], 47[0], 49[1], 70[0], 97[1], 98[1]*, 99[1], and 145[1]. Node K: 86[1], 134[1], and 143[1]. Node L: 31[1], 33[1], 113[1], 116[1], 122[0], 141[1], and 146[1]. Node M: 36[1], 63[0], 65[0], 86[0], 104[1], 117[2], 132[3], 143[0], 136[1], 150[0], and 152[0]. Node M1: 25[0], 51[1], 56[0], 128[1], 133[1], 135[1], and 152[2]. Node M2: 31[0], 38[1], 41[1], 45[1], 49[2], 50[0], 59[1], 75[1], 91[1], 93[1], 112[0], 118[0], 123[0], 129[2], and 148[0].

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Figure 16 in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 16. Ascalabos voithii. Restoration in lateral view, updated from Arratia (1997).

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Figure 8. Ebertichthys ettlingensis n. gen. et n in New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships

Figure 8. Ebertichthys ettlingensis n. gen. et n. sp. Detail of vertebrae 21–23 and associated intermuscular bones (JME ETT 132a). Small arrows point to a median bone that is bifid distally. Abbreviations: Ant, anterior; epin.p, epineural processes; epl.b, epipleural bones; ha, haemal arch; hs, haemal spine; na, neural arch; ns, neural spine; V21, vertebra 21.

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