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FIGURE 10 in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 10. Sketch map of Iran with marked distribution of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species. Red square represents the type locality.

FIGURE 9 in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová & Král, new species (red). Squares represents the type locality of each species.

FIGURE 7A–C in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 7A–C. Holotype of Bolbelasmus nireus (Reitter, 1895). A, habitus in dorsal view; B, original labels of holotype; C, habitus in lateral view. Not to scale.

FIGURE 6A–C in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 6A–C. Allotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species. A, habitus in dorsal view; B, habitus in lateral view; C, pronotum in dorsal view. Not to scale.

FIGURE 4A–D in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 4A–D. Pronotum of males in lateral view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species, from Fars province, Iran; B, B. zagrosensis Sommer, Hillert, Hrůzová & Král, new species, from Lorestan province, Iran; C, B. makrisi Miessen, 2011; D, B. nireus (Reitter, 1895) from Greece, Rhodes. Not to scale.

FIGURE 1A–D in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 1A–D. Habitus of males in dorsal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 2A–D in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 2A–D. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 5A–C in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 5A–C. Parameres in dorsal view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895). Not to scale.

FIGURE 3A–D in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 3A–D. Habitus of males in lateral view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 8A–B in Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus

FIGURE 8A–B. Holotype of Bolbelasmus nireus (Reitter, 1895). A, head in dorsal view; B, pronotum in frontal view. Not to scale.

FIG. 16 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 16. BolbaOEer bidenticollis male: abdomen, posterior view (a), posterior part of abdomen, ventral view (b), aedeagus, dorsal (c), ventral (d), lateral view (e); B. bidenticollis female: abdomen, posterior view (f).

FIG. 17 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 17. BolbaOEer nikolajevi male: abdomen, posterior view (a), posterior part of abdomen, ventral view (b), aedeagus, dorsal (c), ventral (d), lateral view (e); B. nikolajevi female: abdomen, posterior view (f), posterior part of abdomen, ventral view (g).

FIG. 13 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 13. BolbaOEer coriaceus, northern populations: aedeagus with everted internal sac, dorsal (a), ventral (b), lateral view (c); B. coriaceus, southern populations: aedeagus, dorsal (d), ventral (e), lateral (f); B. dudleyi: aedeagus, dorsal (g), ventral (h), lateral view (i); B. abditus: aedeagus, dorsal (j), ventral (k), lateral view (l).

FIG. 8 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 8. BolbaOEer bremeri: aedeagus, dorsal (a), ventral (b), lateral view (c); B. splendidus: aedeagus, dorsal (d), ventral (e), lateral (f), dorsal view with everted internal sac (g); B. bremeri female: posterior part of abdomen, ventral view (h); B. splendidus female: posterior part of abdomen, ventral (i), lateral view (j).

FIG. 12 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 12. Protibial spur, dorsal view: BolbaOEer coriaceus male (a), B. dudleyi male (h), B. dudleyi male and B. abditus male (m); metatrochanter, ventral view: B. tenuelimbatus male and female (b), B. coriaceus male, northern populations (c), southern populations (d), female (e), B. dudleyi male (i), female (j), B. abditus male (n), female (o); B. coriaceus male: abdomen, posterior view (f), posterior part of abdomen, ventral view (g); B. dudleyi male: abdomen, posterior view (k), posterior part of abdomen, ventral view (l); B. abditus male: abdomen, posterior view (p), posterior part of abdomen, ventral view (q).

FIG. 10 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 10. BolbaOEer barbatus, male holotype: metatrochanter, ventral view (a), posterior part of abdomen, ventral view (b); aedeagus with everted internal sac, dorsal (c), ventral (d), lateral view (e).

FIG. 9 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 9. BolbaOEer tenuelimbatus male: abdomen, posterior view (a), posterior part of abdomen, ventral view (b); aedeagus, dorsal (c), ventral (d), lateral view (e); B. tenuelimbatus female: abdomen, posterior view (f).

FIG. 4 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 4. Pronotal armature, frontal view: BolbaOEer princeps major male (a), intermediate male (b), minor male (c), major male (d), intermediate male (e), female (f); B. gigas major male (g), intermediate male (h), minor male (i), female (j); B. mozambiquensis major male (k); B. sasakii major (l), minor male (m), female (n); B. sebastiani major male (o), minor male (p), female (q); B. namibiensis major male (r), intermediate male (s), female (t).

FIG. 5 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 5. Pronotal armature, frontal view: BolbaOEer abyssinicus major male (a), intermediate male (b), minor male (c), female (d).

FIG. 1 in Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species

FIG. 1. BolbaOEer spp.: metasternum (a); posterior end of prosternum, frontal (b), lateral view (c); scutellum and strial punctures (d); metatibia, dorsal view (e); metatibia, lateral view (f). Scale bars in all ®gures 5 1 mm.