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Fig. 11 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 11. Native vegetation (pine forest and montane shrubland) at the top of Cerro del Águila, Mixteca, Oaxaca.

Fig. 10 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 10. Projection of potential distribution for Geotrupes viridiobscurus Jekel, 1866 and G. pecki Howden, 1974 in the state of Oaxaca, representing

Fig. 9. Geotrupes viridiobscurus Jekel, 1866 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 9. Geotrupes viridiobscurus Jekel, 1866: a – male dorsal view (17.44361°N, -96.50361°W); b – female dorsal view (16.160961°N, -96.244523°W); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Fig. 8. Geotrupes pecki Howden, 1974 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 8. Geotrupes pecki Howden, 1974: a – male dorsal view (16.17325°N, -96.24708°W); b – female dorsal view (16.12138°N, -96.45222°W); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Fig. 5 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 5. Map showing the known distribution of Geotrupes lobatus Howden, 1974, G. sallei Jekel, 1866, and G. nebularum Howden, 1974 (upper map) and the known distribution of G. nuntiatus Kohlmann & Arriaga-Jiménez, sp. nov., G. viridiobscurus Jekel, 1866, and G. pecki Howden, 1974 (lower map).

Fig. 7. Geotrupes nebularum Howden, 1974 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 7. Geotrupes nebularum Howden, 1974: a – male dorsal view (19.03638°N, -97.17138°W); b – female dorsal view (19.2533°N, -97.98433°W); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Fig. 4 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 4. Comparison of male genitalia: a – Ceratotrupes bolivari Halffter & Martínez, 1962; and b – Geotrupes lobatus Howden, 1974. Scale bar = 2 mm.

Fig. 3 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 3. Morphological detail of male protibiae: a – Geotrupes nuntiatus Kohlmann & Arriaga-Jiménez, sp. nov.; b – G. lobatus Howden, 1974; arrow pointing at the apical tooth on the profemur. Scale bar = 2 mm.

Fig. 2. Geotrupes lobatus Howden, 1974 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 2. Geotrupes lobatus Howden, 1974: a – male dorsal view (16.16096°N, -96.24452°W); b – female dorsal view (16.16096°N, -96.24452°W); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Fig. 1 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 1. Geotrupes nuntiatus Kohlmann & Arriaga-Jiménez, sp. nov.: a – male dorsal view (17.13087°N, -97.66402°W); b – female dorsal view (17.129932°N, -97.663673°W); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Fig. 6. Geotrupes sallei Jekel, 1866 in A predicted new Geotrupes from the mountains of Oaxaca, Mexico, and a description of the male of Geotrupes lobatus (Coleoptera: Geotrupidae)

Fig. 6. Geotrupes sallei Jekel, 1866: a – male dorsal view (16.19231°N, -96.45711°W); b – female dorsal view (holotype Howden CMN); c–f – male genitalia: c – in dorsal view, d – in ventral view; e–f – lateral views. Scale bars = 5 mm (a–b), 2 mm (c–f).

Figs 13–24 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Figs 13–24. Head, lateral aspect. Maximally developed males: 13 – Lethrus (Lethrus) halkidikensis sp. nov. (holotype); 16 – L. (L.) perun sp. nov. (holotype); 19 – L. (L.) raymondi Reitter, 1890 (Greece: Néa Filadélfia – OHCB); 22 – L. (L.) strymonensis sp. nov. (holotype). Medium developed males: 14 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 17 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 20 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 23 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Underdeveloped males: 15 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 18 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 21 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 24 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Lettering: a – lateral longitudinal keel on base of ventral mandibular process, b – subapical tooth of ventral mandibular process, c – apical tooth of ventral mandibular process. Schematically, not to scale.

Figs 1–12 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Figs 1–12. Head, frontal aspect. Maximally developed males: 1 – Lethrus (Lethrus) halkidikensis sp. nov. (holotype); 4 – L. (L.) perun sp. nov. (holotype); 7 – L. (L.) raymondi Reitter, 1890 (Greece: Néa Filadélfia – OHCB); 10 – L. (L.) strymonensis sp. nov. (holotype). Medium developed males: 2 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 5 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 8 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 11 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Underdeveloped males: 3 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 6 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 9 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 12 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Lettering: b – subapical tooth of ventral mandibular process, c – apical tooth of ventral mandibular process, d – inferiobasal tooth of ventral mandibular process, e – apical emargination of ventral mandibular process. Schematically, not to scale.

Fig. 45 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Fig. 45. Sketch map of the Vardar / Axiós river basin (Macedonia, Greece) with marked distribution of Lethrus (L.) raymondi Reitter, 1890 empty dot represents the type locality.

Figs 39–42 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Figs 39–42. Habitus in frontolateral aspect. 39 – Lethrus (Lethrus) halkidikiensis sp. nov. (holotype), 40 – L. (L.) perun sp. nov. (holotype), 41 – L. (L.) raymondi Reitter, 1890 (Greece: Néa Filadélfia – OHCB, body length: 28 mm), 42 – L. (L.) strymonensis sp. nov. (holotype).

Figs 25–36 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Figs 25–36. Head, dorsal aspect. Maximally developed males: 25 – Lethrus (Lethrus) halkidikensis sp. nov. (holotype); 28 – L. (L.) perun sp. nov. (holotype); 31 – L. (L.) raymondi Reitter, 1890 (Greece: Néa Filadélfia – OHCB); 34 – L. (L.) strymonensis sp. nov. (holotype). Medium developed males: 26 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 29 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 32 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 35 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Underdeveloped males: 27 – Lethrus (L.) halkidikensis sp. nov. (paratype, Greece: Tagarádes – OHCB); 30 – L. (L.) perun sp. nov. (paratype, Bulgaria: Khotovo – OHCB); 33 – L. (L.) raymondi (Greece: Néa Filadélfia – OHCB); 36 – L. (L.) strymonensis sp. nov. (paratype, Greece: Séres – OHCB). Lettering: a – lateral longitudinal keel on base of ventral mandibular process. Schematically, not to scale.

Fig. 47 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Fig. 47. Type locality of Lethrus (L.) halkidikiensis sp. nov. (Greece: Tagarádes, April 2003; photo David Král).

Fig. 50 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Fig. 50. Type locality of Lethrus (L.) strymonensis sp. nov. (Greece: Séres, April 2003; photo David Král).

Fig. 48 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Fig. 48. Type locality of Lethrus (L.) perun sp. nov. (Bulgaria: Khotovo, April 2006; photo David Král).

Fig. 43 in Three new Lethrus species close to L. raymondi (Coleoptera: Geotrupidae) from the Balkan Peninsula

Fig. 43. Sketch map of the Khalkidiki Peninsula (Greece) with marked distribution of Lethrus (L.) halkidikiensis sp. nov., empty dot represents the type locality.