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Fig. 7 in First palaeoscolecid from the Cambrian (Drumian, Miaolingian) Marjum Formation of western Utah, USA

Fig. 7. Palaeogeographic distribution of Cambrian palaeoscolecids in Laurentia. For each occurrence, the numbers in parentheses refer to the number of palaeoscolecid specimens known (left) and the number of species they represent (right). The Rosella Formation Lagerstätte is positioned between the Lagerstätten of the American Great Basin and those of the Canadian Rockies, because the Cassiar terrane was located near present-day Idaho in the Cambrian, and subsequently migrated more than 2000 km to the North (Pope and Sears 1997). Abbreviations: Fm., Formation; Sh., Shale. Background map adapted from Cocks and Torsvik (2011).

opencc-by-4.0Sep 2021View details →
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Fig. 5 in First palaeoscolecid from the Cambrian (Drumian, Miaolingian) Marjum Formation of western Utah, USA

Fig. 5. Scleritome of the palaeoscolecid Arrakiscolex aasei gen et sp. nov. from the Drumian, Cambrian Marjum Formation in the House Range of Utah, USA (Grey Marjum locality). A. SEM micrographs of paratype (KUMIP 314101). A1, dense and homogeneous distribution of the plates on the surfaces of annuli; A2, interannular region showing alternation between plate-free areas and plate-covered areas ("columns"; arrowheads). B. SEM micrographs of holotype (KUMIP 314100). B1, homogeneous distribution of the plates; B2, single plate, showing the swollen inner region surrounded by a raised marginal rim.

opencc-by-4.0Sep 2021View details →
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Fig. 3 in First palaeoscolecid from the Cambrian (Drumian, Miaolingian) Marjum Formation of western Utah, USA

Fig. 3. SEM micrograph and SEM-EDS elemental maps of the posterior region of KUMIP 204770 (Ottoia? sp.), from the Drumian, Cambrian Marjum Formation in the House Range of Utah, USA (Sponge Gully locality). A1, SEM micrograph of the analyzed area (posterior trunk region); A2–A6, SEM-EDS elemental maps of P, Ca, C, Fe, and Mg, respectively.

opencc-by-4.0Sep 2021View details →
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Fig. 4 in First palaeoscolecid from the Cambrian (Drumian, Miaolingian) Marjum Formation of western Utah, USA

Fig. 4. Palaeoscolecid Arrakiscolex aasei gen et sp. nov. from the Drumian, Cambrian Marjum Formation in the House Range of Utah, USA (Grey Marjum locality). A. Holotype (KUMIP 314100). A1, general view of counterpart with location of close-up; A2, detailed view of trunk annulation; A3, general view of part. B. Paratype (KUMIP 314101), general view, showing trunk annulation and location of SEM micrograph in Fig. 5A1 (dashed line rectangle, arrowheads point to the limits of poorly preserved part of the fossil).

opencc-by-4.0Sep 2021View details →
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Fig. 2 in First palaeoscolecid from the Cambrian (Drumian, Miaolingian) Marjum Formation of western Utah, USA

Fig. 2. Ottoid scalidophoran Ottoia? sp. (KUMIP 204770) from the Drumian, Cambrian Marjum Formation in the House Range of Utah, USA (Sponge Gully locality). A1, general view showing body orientation and location of close-ups; A2, detailed view of anterior region, showing the partially everted introvert and digestive tract; A3, detailed view of posterior region, showing location of possible bases of caudal hooks; A4, SEM micrograph showing trunk annulation, arrowheads point to fossil margins; A5, SEM micrograph of anterior region, showing introvert and a few introvert hooks, arrowheads point to fossil margins; A6, SEM micrograph of introvert hooks.

opencc-by-4.0Sep 2021View details →
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FIGURE 8 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 8. Matrix characters for phylogenetic analysis shown on an unspecified species of dolichometopid trilobite (Glossopleura sp.). First number is character number (see Appendix for descriptions and matrix); number in parentheses is character state demonstrated by illustrated form. (Drawing courtesy of Sam Gon III.)

opencc-by-4.0Dec 2022View details →
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FIGURE 3 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 3. Relative eye position and length for dolichometopids in this study, by taxon and formation group (most traditional Glossopleura species blended). This version of the data shows the intermixing of Glossopleura and traditional Anoria and Sonorapis. Note that within taxa, eye position may vary greatly. Only eye length shows possible distinction between genera. Anoria tontoensis from the Bright Angel Formation (along with one specimen of A. bessus from the Gordon Shale) is smaller-eyed and separates from Glossopleura (including G. mckeei) but only at an arbitrary EL/GL of 0.380. A. lodensis, G. mohavensis, and S. californica all plot within Glossopleura.

opencc-by-4.0Dec 2022View details →
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FIGURE 2 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 2. Relative eye position and length for dolichometopids measured in this study, showing low degree to which the studied specimens demonstrate large, posteriorly positioned eyes (toward upper right; historically features attributed to Glossopleura) or smaller, more anteriorly positioned eyes (toward lower left, normally attributed to Anoria and Sonoraspis). Data set equation shows a shallow slope and very weak correlation.

opencc-by-4.0Dec 2022View details →
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FIGURE 9 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 9. Phylogenetic analysis of 12 dolichometopid taxa for which there are relatively complete specimens, plus two outgroup taxa. A) Strict consensus of 40 MPTs of 41 steps. B) Strict consensus of 4 MPTs of 38 steps, after removal of the less well represented taxon G. bion. C) Strict consensus of 2 MPTs of 35 steps with just 8 dolichometopid taxa, after assuming the following synonymies: G. mckeei = G. boccar; S. californica = G. mohavensis; and G. producta = G. boccar. See text.

opencc-by-4.0Dec 2022View details →
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FIGURE 4 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 4. Comparison of cranial measurements (in mm) in dolichometopids assigned to Glossopleura (blue), Sonoraspis (red), and Anoria (black). A) Eye length relative to glabella length. B) Eye position as represented by mid-ocular distance versus glabella length. Genera coded by color as noted above. Species denoted by symbols as listed alphabetically here: Glossopleura bion (filled blue squares), G. boccar (open blue diamonds), G. campbelli (open blue circles), G. gigantea (open blue ovals), G. lodensis (blue stars), G. mckeei (open blue triangles), G. mohavensis (blue dashes), G. producta (filled blue triangles), G. similaris (open inverted blue triangles), G. stephenensis (filled inverted blue triangles), G. utahensis (vertical blue bars), G. walcotti (filled blue diamonds), G. yatesi (filled blue circles); Sonoraspis californica (filled red squares), S. gomezi (open red squares), S. nelsoni (red pluses); Anoria baton (black pluses), A. bessus (black X), and A. tontoensis (filled black circles),

opencc-by-4.0Dec 2022View details →
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FIGURE 11 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 11. Some representative dolichometopids from other formations included in this study. A–B and D–E) Glossopleura boccar. C) Glossopleura similaris(?) F) Anoria tontoensis. G–H) Glossopleura mohavensis. A) USNM 62703, type of Glossopleura boccar, from the Stephen Formation. B) USNM 62702, specimen originally assigned to G. stephenensis, from the Stephen Formation. C) KUMIP 314054, G. similaris(?), from the Spence Shale (Langston Formation). D) SGDS 1975, G. boccar, from the Chisholm Formation. E) USNM 62714, specimen originally assigned to G. mckeei, from the Bright Angel Formation. F) USNM 62685, Anoria tontoensis, from the Bright Angel Formation. G) LACMIP 10782, specimen originally assigned to "Sonoraspis nelsoni", from the Monola Formation. H) LACMIP 2469/24444.1, specimen originally assigned to "Sonoraspis californica", from the Pole Canyon Formation. (Image in D courtesy of Andrew R.C. Milner. G and H courtesy LACM: Hendy, A., Walker, L., Mertz, W. [2020]. LACM Invertebrate Paleontology. Version 1.9. Natural History Museum of Los Angeles County. Occurrence dataset https://doi.org/ 10.15468/6nxzen accessed via GBIF.org on 2021-03-23.) All scale bars equal 1 cm.

opencc-by-4.0Dec 2022View details →
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FIGURE 10. Dolichometopid trilobite specimens from the Cadiz Formation, California. A in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 10. Dolichometopid trilobite specimens from the Cadiz Formation, California. A) Glossopleura lodensis (USNM 78400a), note eight thoracic segments, elongate T5 pleural spines, anteriorly expanded glabella, and long eyes (impression, lighting from left). B) Glossopleura mohavensis (USNM 78400b), note eight thoracic segments (dorsal exoskeleton, lighting from left). C) Glossopleura mohavensis (LACM 10785; "Sonoraspis californica"), note wide glabella, eight thoracic segments, and axial spine bases (impression, lighting from left). D) Glossopleura mohavensis (LACM 10786; "Sonoraspis californica"), note wide glabella, eight thoracic segments, and axial spine bases (impression, lighting from left). E) Glossopleura mohavensis (MWC 7769), note eight thoracic segments (dorsal exoskeleton, lighting from left). F) Glossopleura mohavensis (MWC 7779), (dorsal exoskeleton, lighting from left). A and B from the Bristol Mountains; C and D from near Section A of Foster (1994) (type section; north of Route 66), Marble Mountains; E and F from UCR 7359, near Section B of Foster (1994) (south of Route 66), Marble Mountains. All scale bars = 1 cm.

opencc-by-4.0Dec 2022View details →
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FIGURE 6 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 6. Anterior expansion of glabella in dolichometopids. A) Comparing AGW/MGW and AGW/PGW, showing Anoria and Sonoraspis less expanded but well mixed with Glossopleura specimens. B) Full data set statistics.

opencc-by-4.0Dec 2022View details →
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FIGURE 1 in Continuous character variation within the Glossopleura-Anoria-Sonoraspis plexus: Dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California

FIGURE 1. Measurements taken of dolichometopid cephala for this study. Abbreviations: AGW, anterior glabella width; CW, cephalon width; EL, eye length; GL, glabella length; MGW, mid-glabella width; MOD, midocular distance; PGW, posterior glabella width.

opencc-by-4.0Dec 2022View details →
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Figure 3 in First record of the chancelloriid Allonnia from the middle Cambrian Wheeler Formation (Drumian, Miaolingian) of western Utah

Figure 3. Details of Allonnia cf. Al. tintinopsis, FHPR 11423, from the Wheeler Formation of western Utah. A) SEM image of several sclerites, showing abapical and adapical horizontal rays and absence of central or principal rays, confirming 3+0 arrangement. B) SEM image of sclerite central in A, showing apparent articulatory facets on a displaced adapical horizontal ray between abapical horizontal rays of presumably the same sclerite. C) SEM image of oval-shaped area of high carbon content (dark gray to black) near transition between orange and dark gray preservation areas of the scleritome. D) SEM image of more apically-placed dark gray preservation area of scleritome showing apparent very elongate rays but relatively low elemental contrast between the fossil and matrix. E) SEM image of the basal part of the scleritome showing very elongate sclerite rays preserved as impressions in the matrix rather than by elemental contrast. F) View of scleritome with areas of details A–E indicated. Scale bar = 5 mm. Scale bars for A–E in each image. Abbreviations: abhr, abapical horizontal ray; adhr, adapical horizontal ray; af, articulatory facet; er, very elongate ray.

opencc-by-4.0Aug 2022View details →
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Figure 1 in First record of the chancelloriid Allonnia from the middle Cambrian Wheeler Formation (Drumian, Miaolingian) of western Utah

Figure 1. Location of the Red Wheeler locality (RAR 116, pink pin) in Utah's West Desert, west of Delta, Utah (A); within Utah (B); and in the western United States (C). All base maps from Google Earth, Map data ©2022 Google). Scales marked in lower left in A and B.

opencc-by-4.0Aug 2022View details →
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Figure 2 in First record of the chancelloriid Allonnia from the middle Cambrian Wheeler Formation (Drumian, Miaolingian) of western Utah

Figure 2. Allonnia cf. Al. tintinopsis, FHPR 11423, from the Wheeler Formation of western Utah. A) Scleritome with detail views in B, C, and D marked with white boxes, showing different preservational areas. Scale = 5 mm. B) Color detail of sclerites from middle of scleritome. Scale = 1 mm. C) SEM view showing contrast between matrix (dark gray) and high-iron sclerites (light). Scale = 1 mm. D) SEM view of basal area of scleritome showing multiple elongate sclerite rays. Scale = 1 mm.

opencc-by-4.0Aug 2022View details →
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Data from: Organic-walled microfossils from the Kistedalen Formation, Norway: acritarch chronostratigraphy of the Baltic Miaolingian and evolutionary trends of placoid acritarchs

<p>New acritarch data from the Kistedalen Formation based on two sections with continuous exposure allow to establish the most detailed Miaolingian acritarch biozonation in Baltica to date. Six interval zones previously established in Gondwana, the <em>Eliasum llaniscum, Cristallinium cambriense, Adara alea, Timofeevia lancarae, Cristallinium dubium</em> and <em>Vulcanisphaera turbata</em> zones, are identified, and their correlations within the Acado-Baltic Province examined. The diagnoses of <em>Retisphaeridium, Cristallinium, Retisphaeridium dichamerum, Retisphaeridium lechistanium, Cristallinium dubium</em> and <em>Cristallinium cambriense </em>are emended and the new species <em>Retisphaeridium rugulatum</em> sp. nov. and <em>Retisphaeridium minimum</em> sp. nov. are described. The diagnosis of R<em>etisphaeridium ovillense </em>(Cramer &amp; Díez) Vanguestaine is emended and the species assigned to <em>Dictyotidium ovillense</em> comb. nov. The main evolutionary changes of the placoid acritarchs during the Miaolingian are analysed. Comparison with previous acritarch data from the Kistedalen Formation suggest that different extraction methodologies may result in significant bias in the recovered fossil record.</p>

opencc-zeroJul 2022View details →
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Synchrotron imagery of phosphatized eggs in Waptia cf. W. fieldensis from the middle Cambrian (Miaolingian; Wuliuan) Spence Shale of Utah

<p>Exceptionally preserved fossil eggs and embryos provide critical information regarding paleoembryogenesis, reproductive strategies, and the early ontogeny of early arthropods, but the rarity of preservation of both eggs and egg-bearing organisms in situ limits their use in detailed evo-devo studies. Burgess Shale-type deposits preserve rare instances of egg-bearing arthropods as carbonaceous compressions, however, the eggs are usually poorly preserved with no compelling evidence of embryos. We describe the first record of a brooding specimen of <i>Waptia </i>cf. <i>fieldensis</i> from the Spence Shale, a Cambrian (Wuliuan stage) Burgess Shale-type deposit in northeastern Utah and southeastern Idaho. This is the first record of an egg-bearing arthropod from the Spence Shale and it exhibits two distinct modes of preservation among eggs within the single clutch: carbonization and phosphatization. Unlike the egg-bearing Burgess Shale specimens, many eggs of this Utah specimen are also preserved three-dimensionally. In addition, synchrotron radiation X-ray tomographic microscopy reveals internal distributions of mineral phases, along with potential remnants of the egg membrane and attachment structures but, as in the Burgess Shale, no explicit traces of developing embryos. The distinct modes of preservation highlight the existence of diagenetic microenvironments within some eggs but not in others during fossilization.</p>

opencc-zeroJul 2021View details →
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New soft-bodied panarthropods from diverse Spence Shale (Cambrian; Miaolingian; Wuliuan) depositional environments

<p><span>The Cambrian (Miaolingian; Wuliuan) Spence Shale Lagerstätte of northern Utah and southern Idaho is one of the most diverse Burgess Shale-type deposits of Laurentia. It yields a diverse fauna consisting of abundant biomineralized and locally abundant soft-bodied fossils, along a range of environments from shallow water carbonates to deep shelf dark shales. Panarthropods are the dominant component throughout the deposit, both in time and space, but while the trilobites and agnostoids are abundant, most of the soft-bodied taxa are only known from very few specimens. Additionally, the knowledge of the soft-bodied panarthropods is currently largely limited to locations in the Wellsville Mountains of northeastern Utah. This contribution describes 21 new soft-bodied panarthropods from six locations, including the first occurrences of soft-bodied panarthropods in the High-Creek, Smithfield Creek, Spence Gulch, and Two-Mile Canyon localities. Additionally, we report the presence of bradoriids, <em>Branchiocaris</em> <em>pretiosa</em>, <em>Perspicaris</em>? <em>dilatus</em>, <em>Naraoia</em>? sp., <em>Thelxiope</em> cf. <em>T</em>. <em>palaeothalassia</em>, and <em>Tuzoia</em> <em>guntheri</em> for the first time from the Spence Shale Lagerstätte, the first occurrence reported outside of the Burgess Shale for <em>Thelxiope</em> cf. <em>T</em>. <em>palaeothalassia</em> and the first Wuliuan occurrence of <em>Tuzoia</em> <em>guntheri</em>. We also report on a new hurdiid carapace element and additional specimens of <em>Buccaspinea</em> <em>cooperi</em>?, <em>Dioxycaris</em> <em>argenta</em>, <em>Hurdia</em> sp., and <em>Tuzoia</em> <em>retifera</em>. This new material improves our understanding of the panarthropod fauna of the Spence Shale Lagerstätte and substantially increases our understanding of the distribution of the described taxa in time and space.</span></p>

opencc-zeroMar 2023View details →

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