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Fig. 11. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 11. Nemastoma bidentatum Roewer, 1914 complex, ♀♀; Rec sem (lateral view, right portion, except F left figure, which is in dorsal view). A. The frame encloses the ovipositor portion with Rec sem. Note: Female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

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Fig. 10. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 10. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Pa-Fe (A–B first image, C, D–E first image, F–G first image; H–J, L–M right, N–O: lateral view; the others in dorso-medial view to show the armament).

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Fig. 6. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 6. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Synchroscopic images of the male Ch basal article and Pa-Fe (medial views, except G, right figure, which is in dorso-medial view).

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Fig. 9. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 9. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Variation of the Ch basal article (medial view) in species, subspecies and hybrids.

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Fig. 2 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 2. The complete distributional area of the Nemastoma bidentatum Roewer, 1914 complex, according to Gruber & Martens (1968), Martens (2006), Schönhofer & Holle (2007) and own data, with marked positions of the type localities of the taxa.

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Fig. 13. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 13. Nemastoma bidentatum Roewer, 1914 complex, ♀♀. From left to right: Silhouettes of the Ch basal article, Pa (both medial view) and Rec sem. Arrows indicate the most identifying characters. Note: female N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

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Fig. 12. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 12. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. From left to right: Ch basal article (medial, lateral views), Pa (medial view) and Pe tip (dorsal, lateral views). Arrows indicate the most identifying characters. Note: Pe of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

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Fig. 8. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 8. Nemastoma bidentatum Roewer, 1914 complex. Glans (dorsal and lateral views). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

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Fig. 7. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 7. Nemastoma bidentatum Roewer, 1914 complex. Penes (lateral view). Note: The penis of N. pluridentatum (Hadži, 1973) stat. nov. is unknown.

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Table 6 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 6.</b> Length of appendage-segments in millimeters (trochanters not presented) in <i>N. relictum</i> Gruber &amp; Martens, 1968 stat. nov. from the vicinity of Tappenkarsee, Obertauern, Austria: &male; holotype (&female;) (from Gruber &amp; Martens 1968).</p><table><tbody><tr><th></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Leg I</b></th><td>0.65 (0.75)</td><td>0.30 (0.30)</td><td>0.50 (0.50)</td><td>0.85 (0.90)</td><td>0.80 (0.85)</td><td>3.10 (3.30)</td></tr><tr><th><b>Leg II</b></th><td>1.10 (1.15)</td><td>0.40 (0.40)</td><td>0.95 (0.90)</td><td>1.70 (1.65)</td><td>1.30 (1.30)</td><td>5.45 (5.40)</td></tr><tr><th><b>Leg III</b></th><td>0.70 (0.75)</td><td>0.30 (0.30)</td><td>0.50 (0.55)</td><td>1.00 (1.00)</td><td>0.75 (0.80)</td><td>3.25 (3.40)</td></tr><tr><th><b>Leg IV</b></th><td>1.10 (1.15)</td><td>0.35 (0.30)</td><td>0.75 (0.75)</td><td>1.50 (1.50)</td><td>1.00 (1.00)</td><td>4.70 (4.70)</td></tr></tbody></table>

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Table 4 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 4.</b> Length of appendage-segments in millimeters (trochanters not presented) in <i>N. bidentatum sparsum</i> Gruber &amp; Martens, 1968 from Sonnenberg, Austria: &male; holotype (&female;) (from Gruber &amp; Martens 1968).</p><table><tbody><tr><th></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Leg I</b></th><td>0.75 (0.75)</td><td>0.30 (0.30)</td><td>0.55 (0.55)</td><td>0.95 (0.95)</td><td>0.80 (0.90)</td><td>3.35 (3.45)</td></tr><tr><th><b>Leg II</b></th><td>1.30 (1.25)</td><td>0.35 (0.40)</td><td>1.00 (1.00)</td><td>1.95 (1.95)</td><td>1.45 (1.45)</td><td>6.05 (6.05)</td></tr><tr><th><b>Leg III</b></th><td>0.75 (0.75)</td><td>0.30 (0.30)</td><td>0.50 (0.50)</td><td>1.05 (1.10)</td><td>0.60 (0.75)</td><td>3.20 (3.40)</td></tr><tr><th><b>Leg IV</b></th><td>1.20 (1.20)</td><td>0.35 (0.30)</td><td>0.75 (0.75)</td><td>1.70 (1.70)</td><td>1.05 (1.00)</td><td>5.05 (4.95)</td></tr></tbody></table>

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Table 5 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 5.</b> Length of appendage-segments in millimeters in <i>N. relictum</i> Gruber &amp; Martens, 1968 stat. nov. from Grosses Kar, Koralpe, Austria: &male; (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.25 (0.25)</td><td>0.65 (0.63)</td><td>0.51 (0.47)</td><td>0.41 (0.37)</td><td>&ndash;</td><td>0.32 (0.25)</td><td>2.14 (1.97)</td></tr><tr><th><b>Leg I</b></th><td>0.21 (0.23)</td><td>0.78 (0.74)</td><td>0.34 (0.40)</td><td>0.63 (0.52)</td><td>0.90 (0.90)</td><td>0.89 (0.76)</td><td>3.75 (3.55)</td></tr><tr><th><b>Leg II</b></th><td>0.20 (0.23)</td><td>1.23 (1.23)</td><td>0.42 (0.39)</td><td>0.98 (0.98)</td><td>1.82 (1.62)</td><td>1.21 (1.35)</td><td>5.86 (5.80)</td></tr><tr><th><b>Leg III</b></th><td>0.20 (0.20)</td><td>0.78 (0.56)</td><td>0.31 (0.31)</td><td>0.56 (0.54)</td><td>1.07 (1.00)</td><td>0.67 (0.80)</td><td>3.59 (3.41)</td></tr><tr><th><b>Leg IV</b></th><td>0.24 (0.20)</td><td>1.13 (1.15)</td><td>0.39 (0.38)</td><td>0.83 (0.77)</td><td>1.56 (1.45)</td><td>1.12 (1.04)</td><td>5.27 (4.99)</td></tr></tbody></table>

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Table 3 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 3.</b> Length of appendage-segments in millimeters in <i>N. bidentatum sparsum</i> Gruber &amp; Martens, 1968 from Prosenjakovci, Slovenia: &male; (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.25 (0.28)</td><td>0.75 (0.74)</td><td>0.52 (0.60)</td><td>0.45 (0.47)</td><td>&ndash;</td><td>0.29 (0.28)</td><td>2.26 (2.37)</td></tr><tr><th><b>Leg I</b></th><td>0.23 (0.24)</td><td>0.68 (0.70)</td><td>0.39 (0.38)</td><td>0.61 (0.64)</td><td>1.06 (1.05)</td><td>0.97 (0.98)</td><td>3.94 (3.99)</td></tr><tr><th><b>Leg II</b></th><td>0.27 (0.31)</td><td>1.30 (1.56)</td><td>0.47 (0.45)</td><td>1.13 (1.01)</td><td>2.03 (1.68)</td><td>0.84 (1.47)</td><td>5.98 (6.48)</td></tr><tr><th><b>Leg III</b></th><td>0.19 (0.23)</td><td>0.87 (0.96)</td><td>0.36 (0.35)</td><td>0.61 (0.64)</td><td>1.15 (1.17)</td><td>0.88 (0.91)</td><td>4.06 (4.26)</td></tr><tr><th><b>Leg IV</b></th><td>0.25 (0.25)</td><td>1.32 (1.21)</td><td>0.39 (0.44)</td><td>0.78 (0.89)</td><td>1.84 (1.84)</td><td>1.18 (1.16)</td><td>5.75 (5.79)</td></tr></tbody></table>

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Table 2 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 2.</b> Length of appendage-segments in millimeters (trochanters not presented) in <i>N. bidentatum bidentatum</i> Roewer, 1914 from B&auml;rental, Austria: &male; (&female;) (from Gruber &amp; Martens 1968).</p><table><tbody><tr><th></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Leg I</b></th><td>0.75 (0.75)</td><td>0.35 (0.35)</td><td>0.55 (0.60)</td><td>1.00 (0.90)</td><td>0.85 (0.85)</td><td>3.50 (3.50)</td></tr><tr><th><b>Leg II</b></th><td>1.20 (0.80)</td><td>0.35 (0.40)</td><td>1.05 (1.05)</td><td>1.90 (1.95)</td><td>1.55 (1.60)</td><td>6.05 (5.80)</td></tr><tr><th><b>Leg III</b></th><td>0.75 (0.85)</td><td>0.30 (0.35)</td><td>0.60 (0.55)</td><td>1.15 (1.05)</td><td>0.80 (0.80)</td><td>3.60 (3.60)</td></tr><tr><th><b>Leg IV</b></th><td>1.25 (1.30)</td><td>0.35 (0.35)</td><td>0.75 (0.75)</td><td>1.75 (1.70)</td><td>1.05 (1.05)</td><td>5.15 (5.15)</td></tr></tbody></table>

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Table 1 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 1.</b> Length of appendage-segments in millimeters in <i>N. bidentatum bidentatum</i> Roewer, 1914 from Ljubelj, Slovenia: &male; (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.25 (0.33)</td><td>0.76 (0.80)</td><td>0.53 (0.56)</td><td>0.46 (0.49)</td><td>&ndash;</td><td>0.29 (0.31)</td><td>2.29 (2.49)</td></tr><tr><th><b>Leg I</b></th><td>0.23 (0.24)</td><td>0.68 (0.70)</td><td>0.39 (0.38)</td><td>0.61 (0.64)</td><td>1.06 (1.05)</td><td>0.97 (0.98)</td><td>3.94 (3.99)</td></tr><tr><th><b>Leg II</b></th><td>0.27 (0.31)</td><td>1.30 (1.56)</td><td>0.47 (0.45)</td><td>1.13 (1.01)</td><td>2.03 (1.68)</td><td>0.84 (1.47)</td><td>5.98 (6.48)</td></tr><tr><th><b>Leg III</b></th><td>0.19 (0.23)</td><td>0.87 (0.96)</td><td>0.36 (0.35)</td><td>0.61 (0.64)</td><td>1.15 (1.17)</td><td>0.88 (0.91)</td><td>4.06 (4.26)</td></tr><tr><th><b>Leg IV</b></th><td>0.25 (0.25)</td><td>1.32 (1.21)</td><td>0.39 (0.44)</td><td>0.78 (0.89)</td><td>1.84 (1.84)</td><td>1.18 (1.16)</td><td>5.75 (5.79)</td></tr></tbody></table>

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Fig. 5. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 5. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Pa (medial view).

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Fig. 4. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

Fig. 4. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Ch basal articles (medial, lateral views).

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Table 12 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 12.</b> Length of appendage-segments in millimeters in <i>Nemastoma bidentatum sneznikensis</i> ssp. nov. from Mt. Sne&zcaron;nik, Slovenia: &male; holotype (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.20 (0.15)</td><td>0.59 (0.58)</td><td>0.43 (0.48)</td><td>0.33 (0.38)</td><td>&ndash;</td><td>0.24 (0.25)</td><td>1.79 (1.84)</td></tr><tr><th><b>Leg I</b></th><td>0.19 (0.18)</td><td>0.73 (0.62)</td><td>0.31 (0.28)</td><td>0.52 (0.51)</td><td>0.79 (0.96)</td><td>0.74 (0.63)</td><td>3.28 (3.18)</td></tr><tr><th><b>Leg II</b></th><td>0.19 (0.23)</td><td>1.16 (0.95)</td><td>0.38 (0.38)</td><td>1.03 (0.93)</td><td>1.46 (1.61)</td><td>1.27 (1.24)</td><td>5.49 (5.34)</td></tr><tr><th><b>Leg III</b></th><td>0.18 (0.17)</td><td>0.70 (0.78)</td><td>0.29 (0.27)</td><td>0.53 (0.52)</td><td>0.86 (1.04)</td><td>0.63 (0.57)</td><td>3.19 (3.35)</td></tr><tr><th><b>Leg IV</b></th><td>0.17 (0.18)</td><td>1.06 (0.98)</td><td>0.33 (0.28)</td><td>0.74 (0.73)</td><td>1.42 (1.44)</td><td>1.01 (0.92)</td><td>4.73 (4.53)</td></tr></tbody></table>

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Table 9 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 9.</b> Length of appendage-segments in millimeters in <i>N. kozari</i> Novak, Kozel, Podlesnik &amp; Raspotnig sp. nov. from Vla&scaron;i&cacute; planina, Bosnia and Herzegovina: &male; holotype (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.25 (0.28)</td><td>0.74 (0.69)</td><td>0.51 (0.55)</td><td>0.41 (0.43)</td><td>&ndash;</td><td>0.28 (0.29)</td><td>2.19 (2.24)</td></tr><tr><th><b>Leg I</b></th><td>0.22 (0.23)</td><td>0.87 (0.76)</td><td>0.35 (0.30)</td><td>0.59 (0.62)</td><td>0.97 (0.91)</td><td>0.82 (0.88)</td><td>3.82 (3.70)</td></tr><tr><th><b>Leg II</b></th><td>0.24 (0.24)</td><td>1.32 (1.22)</td><td>0.44 (0.36)</td><td>1.08 (1.00)</td><td>1.87 (1.72)</td><td>1.46 (1.37)</td><td>6.41 (5.91)</td></tr><tr><th><b>Leg III</b></th><td>0.23 (0.26)</td><td>0.82 (0.78)</td><td>0.33 (0.31)</td><td>0.53 (0.51)</td><td>1.09 (1.06)</td><td>0.82 (0.72)</td><td>3.82 (3.64)</td></tr><tr><th><b>Leg IV</b></th><td>0.27 (0.26)</td><td>1.28 (1.23)</td><td>0.43 (0.38)</td><td>0.88 (0.77)</td><td>1.74 (1.69)</td><td>1.04 (0.79)</td><td>5.64 (5.12)</td></tr></tbody></table>

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Table 10 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence

<p><b>Table 10.</b> Length of appendage-segments in millimeters in <i>N. bidentatum martensi</i> Novak, Slana Novak &amp; Raspotnig ssp. nov. from Poljane pri Podgradu, Slovenia: &male; holotype (&female;).</p><table><tbody><tr><th></th><th><b>Trochanter</b></th><th><b>Femur</b></th><th><b>Patella</b></th><th><b>Tibia</b></th><th><b>Metatarsus</b></th><th><b>Tarsus</b></th><th><b>Total length</b></th></tr></tbody><tbody><tr><th><b>Pedipalp</b></th><td>0.27 (0.26)</td><td>0.73 (0.76)</td><td>0.48 (0.55)</td><td>0.44 (0.47)</td><td>&ndash;</td><td>0.27 (0.29)</td><td>2.19 (2.33)</td></tr><tr><th><b>Leg I</b></th><td>0.23 (0.23)</td><td>0.93 (0.80)</td><td>0.35 (0.35)</td><td>0.60 (0.56)</td><td>1.03 (0.94)</td><td>0.78 (0.89)</td><td>3.92 (3.88)</td></tr><tr><th><b>Leg II</b></th><td>0.22 (0.23)</td><td>1.43 (1.35)</td><td>0.48 (0.45)</td><td>1.15 (1.25)</td><td>1.93 (1.80)</td><td>1.69 (1.65)</td><td>6.88 (6.63)</td></tr><tr><th><b>Leg III</b></th><td>0.24 (0.21)</td><td>0.83 (0.80)</td><td>0.33 (0.33)</td><td>0.60 (0.59)</td><td>1.09 (1.05)</td><td>0.83 (0.83)</td><td>3.93 (3.81)</td></tr><tr><th><b>Leg IV</b></th><td>0.23 (0.21)</td><td>1.33 (1.30)</td><td>0.48 (0.38)</td><td>0.83 (0.83)</td><td>1.59 (1.63)</td><td>0.84 (0.83)</td><td>5.30 (5.18)</td></tr></tbody></table>

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