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Figures 1a–1f in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 1a–1f. Developing from the initial stage to the confronting stage. Figure 1a. Initial stage, showing a defensive posture (left, A1, right, A2). Figure 1b. Initial stage, tentative pinching when mutually touched (left, A1, right, A2). Figure 1c. Initial stage, showing a shielding posture (left, B1, right, B2). Figure 1d. Initial stage, one trying to suppress the other (left, B1, right, B2). Figure 1e. Initial stage, shielding collision (left, B1, right, B2). Figure 1f. Spanning pedipalps, developing into confronting stage (left, A1, right, A2).

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Figure 5 in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figure 5. Duration of each round and that of arm-span competition occurred in each round (in seconds).

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Figures 9a–9f in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 9a–9f. Examples of physical combat in genus Hottentotta Birula, 1908. Figures 9a–9c. Adult males of H. minusalta Vachon, 1959. Figure 9d. An adult pair of H. jayakari (Pocock, 1895). Figure 9e. An adult pair of H. franzwerneri (Birula, 1914). Figure 9f. An adult pair of H. buchariensis (Birula, 1897).

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Figures 10a–10c in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 10a–10c. Examples of three types of behavior observed among several species of Heterometrinae. Figure 10a. Arm-span competition (Chersonesometrus tristis). Figure 10b. Shielding response (strictly defined as an easily triggered response; Heterometrus species do exhibit similar posture but happens only when their pedipalps are suppressed; Srilankametrus yaleensis). Figure 10c. Aggressive response (here shows a pre-posture before "pinching" or "punching"; Heterometrus spinifer).

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Figures 7a–7d in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 7a–7d. Examples of physical combat in other scorpions. Figure 7a. Adult females of Androctonus gonneti Vachon, 1948 aiming at each other with their metasoma (the accompanied juddering behavior cannot be illustrated by the figure). Figure 7b. A pair of adult A. gonneti, the female is controlling the telson of the male. Figures 7c–7d. Adults of Hottentotta salei (Vachon, 1980) controlling the telson of the opponent: female and male (c), and two males (d).

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Figures 3a–3b in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 3a–3b. Comparison of aggression between arm-span competition and stinging fighting behavior. Figure 3a. No fighting occurred after one was pulled over (left, A1, right, C). Figure 3b. Violent fighting occurred at the initial stage when both individuals were infuriated (left, A1, right, B1).

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Figures 8a–8f in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 8a–8f. Examples of physical combat in other scorpions. Figures 8a–8c. Juveniles of Hadrurus arizonensis Ewing, 1928, performing the typical deterrent posture. Figures 8d–8f. Leiurus spp., performing the intimidation behavior: adult males of Leiurus jordanensis Lourenço et al., 2002 and L. haenggii Lowe et al., 2014 (d), adult males of L. jordanensis and L. quinquestriatus (Ehrenberg, 1828) (e), and an adult pair of L. jordanensis (f; male on the left).

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Figures 6a–6d in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 6a–6d. Examples of similar behavior observed between two adult males in other Heterometrinae species. Figure 6a. Heterometrus minotaurus (above) and Heterometrus thorellii (below) (photo: V. Tang). Figure 6b. Chersonesometrus tristis (photo: V. Tang). Figure 6c. H. thorellii (photo: Gentia). Figure 6d. H. thorellii (below) and Heterometrus longimanus (above) (photo: Gentia).

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Figures 2a–2f. Confronting stage. Figure 2a in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figures 2a–2f. Confronting stage. Figure 2a. Spreading the pedipalps, lateral view (left, A2, right, A1). Figure 2b. Spreading the pedipalps, posterior view (distal, A1, proximal, C). Figure 2c. Metasoma of the two males entangling with each other (left, C, right, A1). Figure 2d. One being lifted up by the "metasomal hook" (left, C, right, A1). Figure 2e. Lifting, lateral view (left, A1, right, C). Figure 2f. Lifting, posterior view (proximal, A1, distal, C).

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FIGURE 78. Srilankametrus Couzijn, 1978 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 78. Srilankametrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, ventral aspect. A. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). B. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). C. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. F. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 5 mm.

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FIGURE 77. Srilankametrus Couzijn, 1978 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 77. Srilankametrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, lateral aspect. A. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). B. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). C. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. F. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 5 mm.

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FIGURE 76. Srilankametrus Couzijn, 1978 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 76. Srilankametrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, dorsal aspect. A. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). B. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). C. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. F. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 5 mm

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FIGURE 57. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 57. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E–H. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. I–L. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 2 mm.

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FIGURE 56. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 56. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). E–H. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). I–L. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm.

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FIGURE 42. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 42. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., sternum and pectines, ventral aspect. A, B. Srilankametrus indus (DeGeer, 1778), comb. nov., A. ♂, B. ♀ (AMNH), Kandy, Sri Lanka. C, E, F. Srilankametrus serratus (Pocock, 1900), comb. nov., C. holotype ♂ (BMNH 1888.55), E. ♂ (FMNH 086 916), F. ♀ (ZMH), Ceylon [Sri Lanka]. D. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. Scale bars = 2 mm.

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FIGURE 41. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 41. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., sternum and pectines, ventral aspect. A, B. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., A. ♂, B. ♀ (MNHN RS 0079). C, D. Srilankametrus couzijni, sp. nov., C. holotype ♂ (BMNH 1896.7.30.121), D. paratype ♂ (SMF 5332/1). E, F. Srilankametrus gravimanus (Pocock, 1894), comb. nov., E. ♂, F. ♀ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm.

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FIGURE 27. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 27. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., carapace, dorsal aspect. A, B. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., A. ♂, B. ♀ (MNHN RS 0079). C, D. Srilankametrus couzijni, sp. nov., C. holotype ♂ (BMNH 1896.7.30.121), D. paratype ♂ (SMF 5332/1). E, F. Srilankametrus gravimanus (Pocock, 1894), comb. nov., E. ♂, F. ♀ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm.

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FIGURE 28. Srilankametrus Couzijn, 1981 in Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005

FIGURE 28. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., carapace, dorsal aspect. A, B. Srilankametrus indus (DeGeer, 1778), comb. nov., A. ♂, B. ♀ (AMNH), Kandy, Sri Lanka. C, E, F. Srilankametrus serratus (Pocock, 1900), comb. nov., C. holotype ♂ (BMNH 1888.55), E. ♂ (FMNH 086 916), F. ♀ (ZMH), Ceylon [Sri Lanka]. D. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. Scale bars = 2 mm.

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Figure 11 in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figure 11. Dinorhax rostrumpsittaci (Simon, 1877), two males in threatening condition.

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Figure 4 in Non-aggressive competition between males of Srilankametrus yaleensis (Kovařík et al., 2019) (Scorpionidae), and other types of agonistic behavior observed in scorpions

Figure 4. Escaping stage (right male (A2) fled).

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