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Figure 5 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
Figure 5. Tortanus (Atortus) minicoyensis sp. nov., male (allotype): A) habitus, dorsal view; B, C) antennule; D) urosome right lateral view; E) right leg 5 posterior view; F) left leg 5 posterior view; G) second exopodal segment of left leg 5. Cox, coxa; Bas, basis; Exp, exopod; Exp 1, first exopod; Exp 2, second exopod. Arrows indicate 3 rounded processes on the basis of left leg 5.
Figure 3 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
Figure 3. Tortanus (Atortus) minicoyensis sp. nov., female (holotype): A) antenna; B) mandible palp; C) mandible gnathobase; D) maxillule; E) maxilla; F) maxillipede; G) leg 5.
Figure 4 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
Figure 4. Tortanus (Atortus) minicoyensis sp. nov. female (holotype), legs 1–4 anterior view: A) leg 1; B) leg 2; C) leg 3; D) leg 4.
Figure 2 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
Figure 2. Tortanus (Atortus) minicoyensis sp. nov., female (holotype): A) habitus, dorsal view; B) antennule; C) urosome dorsal view (arrows indicate 2 rounded processes on the caudal rami); D) urosome ventral view; the coupling device is dotted in A, C, D.
Figure 1 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
Figure 1. Map showing sampling locations at Minicoy Island, Union Territory of Lakshadweep, southeastern Arabian Sea.
Fig. 3 in Taxonomic description of the male Tortanus (Atortus) murrayi Scott A., 1909 (Copepoda, Calanoida, Tortanidae) from the Great Nicobar Island, Indian Ocean
Fig. 3 — Tortanus (Atortus) murrayi Scott A., 1909, male (a) habitus dorsal view; (b – d) antennule; (e) right leg anterior view; and (f) left leg 5 anterior view
Fig. 3 in Two species of Tortanus (Eutortanus) (Copepoda: Calanoida: Tortanidae) new to Korea
Fig. 3. Tortanus (Eutortanus) vermiculus, male. A. habitus, lateral view. B. habitus, dorsal view. C. right antennule. D. leg 5, posterior view. E. right leg 5, anterior view. Scale bars: A, B=0.25 mm, C-E=0.1 mm.
Fig. 6 in Two species of Tortanus (Eutortanus) (Copepoda: Calanoida: Tortanidae) new to Korea
Fig. 6. Tortanus (Eutortanus) komachi, male. A-D. variability of prosomal end, dorsal view. E. habitus, dorsal view. F. right antennule. G. leg 5, anterior view. Scale bars: A-D=0.25 mm, E=0.5 mm, F-G=0.1 mm.
Fig. 2 in Two species of Tortanus (Eutortanus) (Copepoda: Calanoida: Tortanidae) new to Korea
Fig. 2. Tortanus (Eutortanus) vermiculus, female. A. leg 1, anterior view. B. leg 2, anterior view. C. leg 3, anterior view. D. leg 4, anterior view. E. leg 5, anterior view. Scale bar: A-E=0.1 mm.
Fig. 5 in Two species of Tortanus (Eutortanus) (Copepoda: Calanoida: Tortanidae) new to Korea
Fig. 5. Tortanus (Eutortanus) komachi, female. A. leg 1, posterior view. B. leg 2, posterior view. C. leg 3, posterior view. D. leg 4, posterior view. E. leg 5, anterior view. Scale bar: A-E=0.1 mm.
Table 4 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
<p><b>Table 4.</b> Differential diagnosis of the female <i>T. minicoyensis</i> <b>sp. nov.</b> from all other species of the subgenus <i>Atortus</i> from the Indian Ocean.</p><table><tbody><tr><th><b>Species</b></th><th><b>Fifth pedigerous somite</b></th><th><b>Coupling device</b></th><th><b>Genital compound somite</b></th><th><b>Caudal rami</b></th></tr></tbody><tbody><tr><th><b><i>T. minicoyensis</i> sp. nov.</b></th><td>Asymmetrical, left margin with a notch and right margin with a single downward directed triangular lobe (Figures 2A, 2C, 2D)</td><td>Left process larger than right covering the Asymmetrical, right dorsolateral process of fifth margin with a bulge pedigerous somite and at midpoint with 2 lateral surface of genital ventrolateral spines compound somite (Figure 2D) (Figures 2C, 2D)</td><td>Asymmetrical with 2 rounded medial processes, left ramus slightly larger than the right (Figures 2C, 2D)</td></tr><tr><th><i>T. andamanensis</i></th><td>The right process produced more posteriorly than the left (Nishida et al., 2015, fig. 1A)</td><td>Right process larger, covering the dorsolateral process of the fifth pediger and right lateral surface of genital compound somite (Nishida et al., 2015, figs. 1A, 1B)</td><td>Right anterolateral margin slightly swollen (Nishida et al., 2015, fig. 1D)</td><td>Symmetrical with left ramus slightly produced medially while the right ramus concave (Nishida et al., 2015, figs. 1A, 1D)</td></tr><tr><th><i>T. magnonyx</i></th><td>Asymmetrical, with right side more swollen. (Ohtsuka and Conway, 2005, fig. 2A)</td><td>Not described</td><td>Asymmetrical, expanded anterolaterally, with a pointed process on the right ventrolateral side (Ohtsuka and Conway 2005, figs. 2A–2H)</td><td>Asymmetrical, left ramus larger and wider than right, constricted at base (Ohtsuka and Conway, 2005, figs. 2A, 2C, 2D)</td></tr><tr><th><i>T. insularis</i></th><td>Symmetrical in dorsal view (Ohstuka and Conway, Not described 2003, figs. A, F)</td><td>Asymmetrical, expanded anterolaterally on both sides with peg sensillum at tip (Ohstuka and Conway, 2003, figs. 2C–2E)</td><td>Asymmetrical, left ramus larger than right (Ohstuka and Conway, 2003, figs. 2A, 2C, 2D)</td></tr><tr><th></th><td>Nearly symmetrical, the</td><td></td><td></td><td>Asymmetrical with</td></tr><tr><th><i>T. tropicus</i></th><td>right side being produced backwards in a wing-like process</td><td>Not described (Sewell, 1932, fig. 131a)</td><td>Asymmetrical, small bulge on the left side (Sewell, 1932, fig. 131a)</td><td>distinctly stouter left ramus at its base than that of right (Sewell, 1932, fig.</td></tr><tr><th></th><td>(Sewell, 1932, fig. 131a)</td><td></td><td></td><td>131a)</td></tr><tr><th></th><td></td><td></td><td>Asymmetrical, expanded</td><td></td></tr><tr><th></th><td></td><td></td><td>anterolaterally on both</td><td></td></tr><tr><th></th><td></td><td></td><td>sides ending in small</td><td>Asymmetrical, left ramus</td></tr><tr><th><i>T. recticauda</i></th><td>Symmetrical (Bowman, 1987, fig. 1A)</td><td>Not described</td><td>papilla with apical spinule; right expansion strongly</td><td>slightly wider and longer than right (Bowman,</td></tr><tr><th></th><td></td><td></td><td>convex to straight laterally</td><td>1987, figs. 1A, 1C)</td></tr><tr><th></th><td></td><td></td><td>(Bowman, 1987, figs.</td><td></td></tr><tr><th></th><td></td><td></td><td>1A–1C)</td><td></td></tr></tbody></table>
Table 5 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
<p><b>Table 5.</b> Differential diagnosis of the male <i>T. minicoyensis</i> <b>sp. nov.</b> from all other species of the subgenus <i>Atortus</i> from Indian Ocean.</p><table><tbody><tr><th><b>Species</b></th><th><b>Serrated ridge of right antennule</b></th><th><b>Right leg 5 coxa</b></th><th><b>Right leg 5 basis</b></th><th><b>Left leg 5 subdistal seta</b></th></tr></tbody><tbody><tr><th></th><td>Produced proximally over</td><td></td><td>Bearing triangular shaped</td><td></td></tr><tr><th><b><i>T. minicoyensis</i> sp. nov.</b></th><td>one-third of segment XIX and the ridge slightly raised from the surface plane of the segment</td><td>Semitrapezoid with a beak-shaped medial process (Figure 5E)</td><td>medial process with depression on distomedial margin and fused at the distal side of its base</td><td>Strongly curved along hemispherical tip with the granular surface (Figures 5F, 5G)</td></tr><tr><th></th><td>(Figures 5B, 5C)</td><td></td><td>(Figure 5E)</td><td></td></tr><tr><th></th><td></td><td></td><td>Semicircular and ocarina-</td><td></td></tr><tr><th></th><td></td><td></td><td>shaped medial process</td><td></td></tr><tr><th><i>T. andamanensis</i></th><td>Produced proximally over one-third of segment XIX (Nishida et al., 2015, fig. 4D)</td><td>Semitrapezoid with semispherical medial process (Nishida et al., 2015, figs 4F, 4G)</td><td>with depression on distomedial margin and with the small rounded process at the distal side</td><td>Blunt distal seta (Nishida et al., 2015, figs. 4F, 4H)</td></tr><tr><th></th><td></td><td></td><td>of its base (Nishida et al.,</td><td></td></tr><tr><th></th><td></td><td></td><td>2015, figs. 4F, 4G)</td><td></td></tr><tr><th><i>T. sigmoides</i></th><td>Sigmoid shape and produced proximally to the middle of segment XIX (Nishida et al., 2015, fig. 5D)</td><td>Semicircular with the small bilobed medial process (Nishida et al., 2015, figs. 5F, 5G)</td><td>Semicircular with the fingerlike medial process (Nishida et al., 2015, figs. 5F, 5G)</td><td>Short subdistal seta (Nishida et al., 2015, figs. 5F, 5H)</td></tr><tr><th></th><td></td><td></td><td>Basis massive, tapering</td><td></td></tr><tr><th><i>T. magnonyx</i></th><td>Not produced over the segment XIX (Ohtsuka and Conway, 2005, fig. 3C)</td><td>Coxa unarmed (Ohtsuka and Conway, 2005, figs. 3D–3F)</td><td>proximally, with triangular process bearing two spiniform setae on (Ohtsuka and Conway,</td><td>Terminal seta serrated along inner margin (Ohtsuka and Conway, 2005; figs. 3D, 3H)</td></tr><tr><th></th><td></td><td></td><td>2005, figs. 3D–3F)</td><td></td></tr><tr><th><i>T. insularis</i></th><td>Massive basis, bearing Unarmed bearing 3 acutely Pointed tip with serrated Not produced over the large inner bilobed process pointed prominences of seta (Ohtsuka and segment XIX (Ohtsuka at midlength (Ohtsuka and unequal size (Ohtsuka and Conway, 2003, figs. 3E, and Conway, 2003, fig. 3D) Conway, 2003, Conway, 2003, fig. 3G) 3H) figs. 3E, 3G)</td></tr><tr><th><i>T. nishidai</i></th><td>Serrated ridge on segment XX extending almost along anterior margin of the segment (Ohtsuka et al., 2000, fig. 3D)</td><td>Coxa produced medially into rectangular process (Ohtsuka et al., 2000, figs. 3E, 3H)</td><td>Basis constricted proximally bearing minute seta at midlength on posterior surface (Ohtsuka et al., 2000; figs. 3E, 3H)</td><td>Spiniform terminal seta (Ohtsuka et al., 2000, figs. 3E–3G)</td></tr><tr><th><i>T. tropicus</i></th><td>Produced completely over the segment XIX (Sewell, 1932, fig. 131f)</td><td>Without any process (Sewell, 1932, fig. 131g)</td><td>Small rounded process (Sewell, 1932, fig. 131g)</td><td>Not clear in the description (Sewell, 1932, fig. 131g)</td></tr><tr><th><i>T. recticauda</i></th><td>Serrated ridge on segment XX slightly extended over segment XIX (Bowman, 1986, fig. 1E)</td><td>Coxa of leg 5 truncate with Basis with bilobed process uncinate corner (Bowman, at midlength (Bowman, 1986, figs. 1G–1J) 1986, fig. 1G–1J)</td><td>Long spine with denticulated margin (Bowman, 1986, figs. 1G, 1H)</td></tr></tbody></table>
Fig. 2 in Taxonomic description of the male Tortanus (Atortus) murrayi Scott A., 1909 (Copepoda, Calanoida, Tortanidae) from the Great Nicobar Island, Indian Ocean
Fig. 2 — Tortanus (Atortus) murrayi Scott A., 1909, male dorsal view
Fig. 1 in Taxonomic description of the male Tortanus (Atortus) murrayi Scott A., 1909 (Copepoda, Calanoida, Tortanidae) from the Great Nicobar Island, Indian Ocean
Fig. 1 — Map showing sampling location at Great Nicobar Island, Andaman & Nicobar archipelago
Table 2 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
<p><b>Table 2.</b> Details of sequences incorporated and species abbreviations used in the molecular analysis, as applied in Table 3.</p><table><tbody><tr><th><b>Species</b></th><th><b>Abbreviation</b></th><th><b>Accession No.</b></th><th><b>Remarks</b></th></tr></tbody><tbody><tr><th><i>Tortanus</i> <b>(</b><i>Atortus</i><b>)</b> <i>minicoyensis</i> <b>sp. nov.</b> ♂</th><td>TM♂</td><td>KP749951–KP749953</td><td>Developed</td></tr><tr><th><i>Tortanus</i> <b>(</b><i>Atortus</i><b>)</b> <i>minicoyensis</i> <b>sp. nov.</b> ♀</th><td>TM♀</td><td>KP749954</td><td>Developed</td></tr><tr><th><i>T.</i> <b>(</b><i>Boreotortanus</i><b>)</b> <i>discaudatus</i></th><td>TD</td><td>AF513648</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Atortus</i><b>)</b> <i>erabuensis</i></th><td>TE</td><td>AF474113</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Tortanus</i><b>)</b> <i>forcipatus</i></th><td>TF</td><td>AF513649, AY145431</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Atortus</i><b>)</b> <i>ryukyuensis</i></th><td>TR</td><td>AF474114</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Atortus</i><b>)</b> <i>insularis</i></th><td>TI</td><td>KC287905</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Eutortanus</i><b>)</b> <i>derjugini</i></th><td>TDE</td><td>AF513647, AF474112</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Eutortanus</i><b>)</b> <i>dextrilobatus</i></th><td>TDX</td><td>AY145430</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Eutortanus</i><b>)</b> <i>komachi</i></th><td>TK</td><td>JN605792</td><td>Obtained</td></tr><tr><th><i>T.</i> <b>(</b><i>Eutortanus</i><b>)</b> <i>vermiculus</i></th><td>TV</td><td>JN605791</td><td>Obtained</td></tr><tr><th><i>Nassodonta insignis</i></th><td>NI</td><td>KT985464</td><td>Obtained</td></tr></tbody></table>
Table 3 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
<p><b>Table 3.</b> Distance matrix showing inter- and intraspecific percentage divergence of <i>Tortanus minicoyensis</i> <b>sp. nov.</b> and other species of the genus <i>Tortanus.</i> See Table 2 for codes for specimens and species.</p><table><tbody><tr><th></th><th>Accession No.</th><th><b>1</b></th><th><b>2</b></th><th><b>3</b></th><th><b>4</b></th><th><b>5</b></th><th><b>6</b></th><th><b>7</b></th><th><b>8</b></th><th><b>9</b></th><th><b>10</b></th><th><b>11</b></th><th><b>12</b></th><th><b>13</b></th><th><b>14</b></th><th><b>15</b></th><th><b>16</b></th></tr></tbody><tbody><tr><th><b>1</b></th><td>KP749952 TM ♂</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>2</b></th><td>KP749953 TM ♂</td><td>0.0</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>3</b></th><td>KP749951 TM ♂</td><td>0.0</td><td>0.0</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>4</b></th><td>KP749954 TM♀</td><td>0.4</td><td>0.4</td><td>0.4</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>5</b></th><td>AF513648 TD</td><td>22.9</td><td>22.9</td><td>22.9</td><td>22.7</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>6</b></th><td>AF474113 TE</td><td>18.7</td><td>18.7</td><td>18.7</td><td>18.7</td><td>16.9</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>7</b></th><td>AF513649 TF</td><td>21.9</td><td>21.9</td><td>21.9</td><td>21.9</td><td>21.2</td><td>19.9</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>8</b></th><td>AY145431TF</td><td>19.2</td><td>19.2</td><td>19.2</td><td>19.2</td><td>19.3</td><td>18.4</td><td>3.3</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>9</b></th><td>KC287905 TI</td><td>20.9</td><td>20.9</td><td>20.9</td><td>20.9</td><td>19.0</td><td>20.9</td><td>22.8</td><td>21.2</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>10</b></th><td>AF513647 TDE</td><td>17.9</td><td>17.9</td><td>17.9</td><td>17.4</td><td>19.6</td><td>15.4</td><td>22.5</td><td>20.9</td><td>21.4</td><td></td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>11</b></th><td>AF474112 TDE</td><td>18.4</td><td>18.4</td><td>18.4</td><td>18.4</td><td>19.0</td><td>15.7</td><td>23.4</td><td>21.5</td><td>20.6</td><td>1.4</td><td></td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>12</b></th><td>AY145430 TDX</td><td>20.3</td><td>20.3</td><td>20.3</td><td>20.3</td><td>19.0</td><td>20.1</td><td>23.6</td><td>22.0</td><td>20.0</td><td>8.6</td><td>8.4</td><td></td><td></td><td></td><td></td><td></td></tr><tr><th><b>13</b></th><td>AF474114 TR</td><td>22.5</td><td>22.5</td><td>22.5</td><td>22.5</td><td>25.3</td><td>20.1</td><td>24.2</td><td>23.7</td><td>23.3</td><td>18.0</td><td>18.5</td><td>20.9</td><td></td><td></td><td></td><td></td></tr><tr><th><b>14</b></th><td>JN605792 TK</td><td>21.0</td><td>21.0</td><td>21.0</td><td>21.0</td><td>18.2</td><td>21.1</td><td>24.2</td><td>23.4</td><td>20.8</td><td>19.1</td><td>19.1</td><td>19.1</td><td>25.5</td><td></td><td></td><td></td></tr><tr><th><b>15</b></th><td>JN605791 TV</td><td>18.4</td><td>18.4</td><td>18.4</td><td>19.0</td><td>18.5</td><td>18.4</td><td>23.6</td><td>21.7</td><td>18.8</td><td>15.4</td><td>14.6</td><td>17.2</td><td>22.5</td><td>19.7</td><td></td><td></td></tr><tr><th><b>16</b></th><td>KT985464 NI</td><td>34.4</td><td>34.4</td><td>34.4</td><td>34.4</td><td>36.0</td><td>34.4</td><td>35.7</td><td>33.1</td><td>34.7</td><td>35.7</td><td>35.0</td><td>34.0</td><td>40.0</td><td>38.7</td><td>34.4</td><td></td></tr></tbody></table>
Table 1 in A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea
<p><b>Table 1</b>. Seta (in Arabic numerals) and spine formula (in Roman numerals) of legs 1–4 of female <i>Tortanus minicoyensis</i> sp. nov.</p><table><tbody><tr><th></th><th><b>Coxa</b></th><th><b>Basis</b></th><th><b>Exopod segment 1; 2; 3</b></th><th><b>Endopod segment 1; 2</b></th></tr></tbody><tbody><tr><th><b>Leg 1</b></th><td>0–1</td><td>1–0</td><td>0–1; 0–1; I, I, 4</td><td>0–3; 1, 2, 3</td></tr><tr><th><b>Leg 2</b></th><td>0–1</td><td>0–0</td><td>I–1; I–1; III, I, 5</td><td>0–3; 1, 2, 3</td></tr><tr><th><b>Leg 3</b></th><td>0–1</td><td>0–0</td><td>I–1; I–1; III, I, 5</td><td>0–3; 1, 2, 3</td></tr><tr><th><b>Leg 4</b></th><td>0–1</td><td>1–0</td><td>I–1; I–1; III, I, 5</td><td>0–3; 1, 2, 3</td></tr></tbody></table>
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Allen Brain Atlas
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