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Text-fig. 3. Pinaceae, Taxaceae. a: Pinus needle fascicle with 3 needles, UAPC-ALTA S 25088A. b: Pinus needle fascicle with at least 4 needles, UAPC-ALTA S 59496. c: Articulate Pinus seed (section Diploxylon) showing seed body partly detached from wing, BBM-PAL-P000007. d: Winged pinaceous seed with elongate, flattened wing and narrow seed body, BBM-PAL-P000048. e: Another winged pinaceous seed with very narrow seed body, BBM-PAL-P000008. f: Amentotaxus leaf, UAPC-ALTA S S25086A. g: Higher magnification counterpart of (f) showing abaxial (lower) leaf surface with two parallel stomatal bands and tapered leaf tip S 25086B. h: Higher magnification of specimen in (f) showing adaxial (upper) leaf surface with detail of single midvein. Scale bars: a–e, g, h = 1 cm, f = 2 cm. in The Early Eocene Flora Of Horsefly, British Columbia, Canada And Its Phytogeographic Significance

Text-fig. 3. Pinaceae, Taxaceae. a: Pinus needle fascicle with 3 needles, UAPC-ALTA S 25088A. b: Pinus needle fascicle with at least 4 needles, UAPC-ALTA S 59496. c: Articulate Pinus seed (section Diploxylon) showing seed body partly detached from wing, BBM-PAL-P000007. d: Winged pinaceous seed with elongate, flattened wing and narrow seed body, BBM-PAL-P000048. e: Another winged pinaceous seed with very narrow seed body, BBM-PAL-P000008. f: Amentotaxus leaf, UAPC-ALTA S S25086A. g: Higher magnification counterpart of (f) showing abaxial (lower) leaf surface with two parallel stomatal bands and tapered leaf tip S 25086B. h: Higher magnification of specimen in (f) showing adaxial (upper) leaf surface with detail of single midvein. Scale bars: a–e, g, h = 1 cm, f = 2 cm.

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Text-fig. 7. Scanning electron micrographs (a, b, d, e, g–k), X-ray microtomographic orthoslices (c) and synchrotron radiation X-ray tomographic microscopy orthoslices (f) of fruits and endocarps of uncertain affinity from Zliv-Řídká Blana locality. a–c: Trebecenia sarcocalis, a – tricarpellate fruit, no. NM-F 3637, b – fruits supported by pentamerous and persistent calyx, no. NMF 3637, c – fruit almost circular in transverse section, no. NM-F 3637; d: Taxon 17, small fruit with slightly sunken stylar region, no. NM-F 3201; e: Taxon 19, spherical fruit, the fruit wall composed of large isodiametric, thick walled cells, no. NM-F 3181; f: Taxon 19, single-seeded fruit, no. NM-F 3621; g: Taxon 20, syncarpous, multicarpellate fruit of ten carpels, no. NM-F 3200; h: Taxon 22, syncarpous, multicarpellate fruit of seven carpels, no. NM-F 3159; i: cf. Sabia menispermoides, endocarp of drupaceous fruits, no. NM-F 4624; j: Taxon 25, endocarp triangular in cross-section, no. NM-F 3218; k: Taxon 24, endocarp spherical in cross-section with a distinctly ribbed and foveolate surface, no. NM-F 3217. in Plant Mesofossils From The Late Cretaceous Klikov Formation, The Czech Republic

Text-fig. 7. Scanning electron micrographs (a, b, d, e, g–k), X-ray microtomographic orthoslices (c) and synchrotron radiation X-ray tomographic microscopy orthoslices (f) of fruits and endocarps of uncertain affinity from Zliv-Řídká Blana locality. a–c: Trebecenia sarcocalis, a – tricarpellate fruit, no. NM-F 3637, b – fruits supported by pentamerous and persistent calyx, no. NMF 3637, c – fruit almost circular in transverse section, no. NM-F 3637; d: Taxon 17, small fruit with slightly sunken stylar region, no. NM-F 3201; e: Taxon 19, spherical fruit, the fruit wall composed of large isodiametric, thick walled cells, no. NM-F 3181; f: Taxon 19, single-seeded fruit, no. NM-F 3621; g: Taxon 20, syncarpous, multicarpellate fruit of ten carpels, no. NM-F 3200; h: Taxon 22, syncarpous, multicarpellate fruit of seven carpels, no. NM-F 3159; i: cf. Sabia menispermoides, endocarp of drupaceous fruits, no. NM-F 4624; j: Taxon 25, endocarp triangular in cross-section, no. NM-F 3218; k: Taxon 24, endocarp spherical in cross-section with a distinctly ribbed and foveolate surface, no. NM-F 3217.

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Text-fig. 6. Scanning electron micrographs of seeds of ericalean affinity (a–d) and seeds of uncertain affinity (e–i) from Zliv-Řídká Blana locality. a: Protovisnea sp. 1, rounded seed with the narrow elongate seed cavity flanked by two bulging regions of larger cells, no. NM-F 3177; b: Protovisnea sp. 2, angular seed with the narrow elongate seed cavity flanked by two bulging regions of larger cells, no. NM-F 3179; c, d: Eurya crassitesta, one seed split into two parts, no. NM-F 3211, c – surface cells of the seed coat are palisade, d – cross-section of the seed; e: Nympheaceae sp. 1, seed, no. NM-F 3636; f: Nympheaceae sp. 2, seed, no. NM-F 4634; g: Klikovispermum sp.1, seeds with irregular outline and smooth outer surface, no. NM-F 3203; h: Klikovispermum malechii, seed with an orange-segment shape, no. NM-F 3299; i: Taxon 35, seed, no. NM-F 3236. in Plant Mesofossils From The Late Cretaceous Klikov Formation, The Czech Republic

Text-fig. 6. Scanning electron micrographs of seeds of ericalean affinity (a–d) and seeds of uncertain affinity (e–i) from Zliv-Řídká Blana locality. a: Protovisnea sp. 1, rounded seed with the narrow elongate seed cavity flanked by two bulging regions of larger cells, no. NM-F 3177; b: Protovisnea sp. 2, angular seed with the narrow elongate seed cavity flanked by two bulging regions of larger cells, no. NM-F 3179; c, d: Eurya crassitesta, one seed split into two parts, no. NM-F 3211, c – surface cells of the seed coat are palisade, d – cross-section of the seed; e: Nympheaceae sp. 1, seed, no. NM-F 3636; f: Nympheaceae sp. 2, seed, no. NM-F 4634; g: Klikovispermum sp.1, seeds with irregular outline and smooth outer surface, no. NM-F 3203; h: Klikovispermum malechii, seed with an orange-segment shape, no. NM-F 3299; i: Taxon 35, seed, no. NM-F 3236.

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Text-fig. 2. SEM images of Miranthus elegans gen. et sp. nov.; Mira locality, Portugal. a, b: Flowers in oblique lateral view showing remains of calyx and slightly semi-inferior ovary with elongated apical style (a); note larger openings in the floral tissue (asterisk) interpreted as schizogenous secretory cavities and the stomata-like secretory structures on the upper portion of the ovary (arrows) that are interpreted as nectariferous (b). c: Detail of ovary surface showing secretory stomata-like structures (arrows). d: Flower in lateral view showing fragmentary calyx and broken slightly semi-inferior ovary with secretory stomata-like structures; note the point of attachment of the central placenta (pl). e: Cluster of seeds removed from the ovary in (d) showing reticulate surface. f: Outer (abaxial) surface of calyx lobe showing the slightly pointed papillae and scattered, fine trichomes (arrows). g: Triaperturate pollen grains from the ovary surface. Specimens, Mira 100-S153146 (a, b), Mira 100-S170155 (c), Mira 100-S101266 (d, e), Mira 105-S100732 (f), Mira 100-S170125 (g). Scale bars = 600 µm (a, b, d), 300 µm (f), 100 µm (c, e), 10 µm (g). in Early Flowers Of Primuloid Ericales From The Late Cretaceous Of Portugal And Their Ecological And Phytogeographic Implications

Text-fig. 2. SEM images of Miranthus elegans gen. et sp. nov.; Mira locality, Portugal. a, b: Flowers in oblique lateral view showing remains of calyx and slightly semi-inferior ovary with elongated apical style (a); note larger openings in the floral tissue (asterisk) interpreted as schizogenous secretory cavities and the stomata-like secretory structures on the upper portion of the ovary (arrows) that are interpreted as nectariferous (b). c: Detail of ovary surface showing secretory stomata-like structures (arrows). d: Flower in lateral view showing fragmentary calyx and broken slightly semi-inferior ovary with secretory stomata-like structures; note the point of attachment of the central placenta (pl). e: Cluster of seeds removed from the ovary in (d) showing reticulate surface. f: Outer (abaxial) surface of calyx lobe showing the slightly pointed papillae and scattered, fine trichomes (arrows). g: Triaperturate pollen grains from the ovary surface. Specimens, Mira 100-S153146 (a, b), Mira 100-S170155 (c), Mira 100-S101266 (d, e), Mira 105-S100732 (f), Mira 100-S170125 (g). Scale bars = 600 µm (a, b, d), 300 µm (f), 100 µm (c, e), 10 µm (g).

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Text-fig. 7. Exbeckettia mastixioides (E.REID et M.CHANDLER) comb. nov. Details of anatomy in transverse section on cut surfaces from bilocular fruit shown in Text-fig. 6i, V. 23013(3). a: Details of seed (S), locule (L), distinct endocarp planes of separation (arrows). b: Detail of endocarp adjacent to the locule, and surrounding mesocarp. Blue lines indicate thickness of sclerenchyma lining the locule. Note layer of horizontally oriented periclinal fibres a few cells thick, lining the locule (arrow). c: Enlargement showing parenchyma cells of the mesocarp decreasing in diameter toward the periphery. d: Enlargement showing fibres and sclereids of the endocarp. e: Sharp contact between endocarp and mesocarp. f, g: Detailed anatomy of endocarp including locule lining, and contact with mesocarp. Scale bars 2 mm in (a), (b), 1 mm in (c–g). in Mastixioid Fruits (Cornales) From The Early Eocene London Clay Flora: Morphology, Anatomy And Nomenclatural Revision

Text-fig. 7. Exbeckettia mastixioides (E.REID et M.CHANDLER) comb. nov. Details of anatomy in transverse section on cut surfaces from bilocular fruit shown in Text-fig. 6i, V. 23013(3). a: Details of seed (S), locule (L), distinct endocarp planes of separation (arrows). b: Detail of endocarp adjacent to the locule, and surrounding mesocarp. Blue lines indicate thickness of sclerenchyma lining the locule. Note layer of horizontally oriented periclinal fibres a few cells thick, lining the locule (arrow). c: Enlargement showing parenchyma cells of the mesocarp decreasing in diameter toward the periphery. d: Enlargement showing fibres and sclereids of the endocarp. e: Sharp contact between endocarp and mesocarp. f, g: Detailed anatomy of endocarp including locule lining, and contact with mesocarp. Scale bars 2 mm in (a), (b), 1 mm in (c–g).

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Text-fig. 2. Salicaceae (a–g), Cannabaceae (h–n), cf. Betulaceae (o–r). a–g: Saxifragispermum, USNM PAL 772341. Scale bar = 5 mm except as indicated. a–b: Lateral, c: apical, and d: basal views of fruit, reflected light, palladium coated; apex at top of (a, b). e: Equatorial transverse section reflected light; arrows indicate presumed seeds, scale bar = 2 mm. f: Detail of locule contents extracted from (e), transmitted light, scale bar = 200 Μm. g: Interwoven trichomes or fibers from locule, transmitted light, scale bar = 5 Μm. h–j: Celtis. h, i: USNM PAL 772342, reflected light, palladium coated, scale bar = 5 mm. h: Lateral view parallel with plane of dehiscence. i: Lateral view perpendicular to plane of dehiscence. j: DMNH EPI.47809, Celtis in lateral view; showing reticulate sculpture and the vertically-oriented, plane of dehiscence (arrow), scale bar = 5 mm. k–m: Aphananthe. USNM PAL 772344, reflected light, palladium coated, scale bar = 5 mm. k: Apical view, note triangular cross section and apical plug (arrow). l: Lateral view, apex up. m: Lateral view at 90° to (l). n: Detail of cellular pattern at surface of endocarp, scale bar = 0.5 mm. o–r: in The Early Middle Eocene Wagon Bed Carpoflora Of Central Wyoming, U.S.A.

Text-fig. 2. Salicaceae (a–g), Cannabaceae (h–n), cf. Betulaceae (o–r). a–g: Saxifragispermum, USNM PAL 772341. Scale bar = 5 mm except as indicated. a–b: Lateral, c: apical, and d: basal views of fruit, reflected light, palladium coated; apex at top of (a, b). e: Equatorial transverse section reflected light; arrows indicate presumed seeds, scale bar = 2 mm. f: Detail of locule contents extracted from (e), transmitted light, scale bar = 200 Μm. g: Interwoven trichomes or fibers from locule, transmitted light, scale bar = 5 Μm. h–j: Celtis. h, i: USNM PAL 772342, reflected light, palladium coated, scale bar = 5 mm. h: Lateral view parallel with plane of dehiscence. i: Lateral view perpendicular to plane of dehiscence. j: DMNH EPI.47809, Celtis in lateral view; showing reticulate sculpture and the vertically-oriented, plane of dehiscence (arrow), scale bar = 5 mm. k–m: Aphananthe. USNM PAL 772344, reflected light, palladium coated, scale bar = 5 mm. k: Apical view, note triangular cross section and apical plug (arrow). l: Lateral view, apex up. m: Lateral view at 90° to (l). n: Detail of cellular pattern at surface of endocarp, scale bar = 0.5 mm. o–r:

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Text-fig. 9. Carpolithes (a–r). a–d: Carpolithes sp. 5. USNM PAL 772370. Scale bar = 5 mm, reflected light, palladium coated. a: Lateral view of seed, apex up, possible raphe descending from apex toward viewer. b: Lateral view of seed, apex up, possible raphe on right. c: Lateral view, opposite side, apex up, possible raphe on left. d: Apical view, note central pit with raphe descending towards bottom margin. e–h: Carpolithes sp. 6. USNM PAL 772371. Scale bar = 5 mm. e: Basal view illustrating depression and keel in plane of bisymmetry, reflected light, palladium coated. f–h: Micro-CT scan surface rendering. f: Lateral view showing relatively smooth rounded surface. g: Specimen rotated 180° from (f), surface partially eroded. h: Longitudinal view, showing median keel. i–m: Carpolithes sp. 7 USNM PAL 772372. Scale bar = 5 mm. i: View of intact face of globose fruit, possible apical constriction at top. j: Lateral view, intact surface to right, possible apical constriction at top, both micro-CT scan surface renderings. k: Apical view. l: Face view illustrating the mineral filling and the fine, radiating structure of the fruit wall on the left and right margins, both reflected light, palladium coated. m: Closeup of the cellular layer on the left of (l), micro-CT scan surface rendering. n–p: Carpolithes sp. 8. USNM PAL 772373. Scale bar = 3 mm, reflected light, palladium coated. n: Lateral view of pyrene-like structure, one ridge running vertically in the center of view, the other two forming the left and right margins. o: Lateral view of pyrene-like structure, ridge in (n) on the left. p: End-on view illustrating one convex, one concave, and one relatively flat to very slightly concave face. q, r: Carpolithes sp. 9 USNM PAL 772374. Scale bar = 5 mm, reflected light, palladium coated. q: Exterior of the smooth broken half-sphere. r: Interior of the broken half-sphere. in The Early Middle Eocene Wagon Bed Carpoflora Of Central Wyoming, U.S.A.

Text-fig. 9. Carpolithes (a–r). a–d: Carpolithes sp. 5. USNM PAL 772370. Scale bar = 5 mm, reflected light, palladium coated. a: Lateral view of seed, apex up, possible raphe descending from apex toward viewer. b: Lateral view of seed, apex up, possible raphe on right. c: Lateral view, opposite side, apex up, possible raphe on left. d: Apical view, note central pit with raphe descending towards bottom margin. e–h: Carpolithes sp. 6. USNM PAL 772371. Scale bar = 5 mm. e: Basal view illustrating depression and keel in plane of bisymmetry, reflected light, palladium coated. f–h: Micro-CT scan surface rendering. f: Lateral view showing relatively smooth rounded surface. g: Specimen rotated 180° from (f), surface partially eroded. h: Longitudinal view, showing median keel. i–m: Carpolithes sp. 7 USNM PAL 772372. Scale bar = 5 mm. i: View of intact face of globose fruit, possible apical constriction at top. j: Lateral view, intact surface to right, possible apical constriction at top, both micro-CT scan surface renderings. k: Apical view. l: Face view illustrating the mineral filling and the fine, radiating structure of the fruit wall on the left and right margins, both reflected light, palladium coated. m: Closeup of the cellular layer on the left of (l), micro-CT scan surface rendering. n–p: Carpolithes sp. 8. USNM PAL 772373. Scale bar = 3 mm, reflected light, palladium coated. n: Lateral view of pyrene-like structure, one ridge running vertically in the center of view, the other two forming the left and right margins. o: Lateral view of pyrene-like structure, ridge in (n) on the left. p: End-on view illustrating one convex, one concave, and one relatively flat to very slightly concave face. q, r: Carpolithes sp. 9 USNM PAL 772374. Scale bar = 5 mm, reflected light, palladium coated. q: Exterior of the smooth broken half-sphere. r: Interior of the broken half-sphere.

Text-fig. 35. Scanning electron microscope (SEM) images of a fruit of Pennicarpus sp. and monocolpate pollen of Pennipollis sp. from a pollen clump; Torres Vedras locality, Portugal. a) Fruit of Pennicarpus sp. showing the poorly preserved fruit wall enclosing a single seed; b) Pollen clump that yielded the pollen in this Text-figure (c, d); c, d) Pollen grains showing the loose reticulum bearing small spinules that are often in two rows; note the smooth surface of the foot layer, the absence of columellae, the continuous muri bordering the apertures and spiny orbicules (arrowheads). Specimens, TV38-S174609 (a), TV44-S148015 (b–d). Scale bars 300 Μm (a, b), 6 Μm (c, d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 35. Scanning electron microscope (SEM) images of a fruit of Pennicarpus sp. and monocolpate pollen of Pennipollis sp. from a pollen clump; Torres Vedras locality, Portugal. a) Fruit of Pennicarpus sp. showing the poorly preserved fruit wall enclosing a single seed; b) Pollen clump that yielded the pollen in this Text-figure (c, d); c, d) Pollen grains showing the loose reticulum bearing small spinules that are often in two rows; note the smooth surface of the foot layer, the absence of columellae, the continuous muri bordering the apertures and spiny orbicules (arrowheads). Specimens, TV38-S174609 (a), TV44-S148015 (b–d). Scale bars 300 Μm (a, b), 6 Μm (c, d).

Text-fig. 24. Scanning electron microscope (SEM) images (a, c, d) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images (b, e) of fruits and seeds of Serialis antiqua (a, b) and Serialis parva (c–e); Torres Vedras locality, Portugal. a, b) Serialis antiqua lateral view of fruit showing smooth partly abraded fruit wall (a) and longitudinal section (b, SRXTM orthoslice yz0862) showing six closely adhering seeds, some of which are mature with well-preserved nutritive tissue; c–e) Serialis parva lateral view of fruit showing faint ribs (c), finely pitted seed surface with undulate anticlinal walls of the exotestal cells (d), and transverse section showing two closely adhering seeds (SRXTM orthoslice xy0913, e). Specimens, TV43-S174480 (a, b), TV43-S170076 (c), TV43-S170079 (d), TV43-S174477 (e). Scale bars 300 Μm (a–c, e), 50 Μm (d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 24. Scanning electron microscope (SEM) images (a, c, d) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images (b, e) of fruits and seeds of Serialis antiqua (a, b) and Serialis parva (c–e); Torres Vedras locality, Portugal. a, b) Serialis antiqua lateral view of fruit showing smooth partly abraded fruit wall (a) and longitudinal section (b, SRXTM orthoslice yz0862) showing six closely adhering seeds, some of which are mature with well-preserved nutritive tissue; c–e) Serialis parva lateral view of fruit showing faint ribs (c), finely pitted seed surface with undulate anticlinal walls of the exotestal cells (d), and transverse section showing two closely adhering seeds (SRXTM orthoslice xy0913, e). Specimens, TV43-S174480 (a, b), TV43-S170076 (c), TV43-S170079 (d), TV43-S174477 (e). Scale bars 300 Μm (a–c, e), 50 Μm (d).

Text-fig. 16. Scanning electron microscope (SEM) images of seeds of possible Nymphaeales-Austrobaileyales affinity; Torres Vedras locality, Portugal. a, b) Nymphaeales-Austrobaileyales seed sp. 1, seed in lateral view (a) showing smooth surface of the exotesta composed of cells with shallowly sinuous cell outlines (b); c, d) Nymphaeales-Austrobaileyales seed sp. 2, fragmentary seed in lateral view (c) showing surface of exotesta composed of cells with deeply sinuous cell outlines (d); e, f) Nymphaeales-Austrobaileyales seed sp. 3, seed in lateral view (e) showing detail of smooth surface of the exotesta composed of cells with sinuous cell outlines (f). Specimens, TV43-S136741 (a, b), TV43-S148142 (c, d), TV43-S170087 (e, f). Scale bars 300 Μm (a, c, e), 30 Μm (b, d, f). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 16. Scanning electron microscope (SEM) images of seeds of possible Nymphaeales-Austrobaileyales affinity; Torres Vedras locality, Portugal. a, b) Nymphaeales-Austrobaileyales seed sp. 1, seed in lateral view (a) showing smooth surface of the exotesta composed of cells with shallowly sinuous cell outlines (b); c, d) Nymphaeales-Austrobaileyales seed sp. 2, fragmentary seed in lateral view (c) showing surface of exotesta composed of cells with deeply sinuous cell outlines (d); e, f) Nymphaeales-Austrobaileyales seed sp. 3, seed in lateral view (e) showing detail of smooth surface of the exotesta composed of cells with sinuous cell outlines (f). Specimens, TV43-S136741 (a, b), TV43-S148142 (c, d), TV43-S170087 (e, f). Scale bars 300 Μm (a, c, e), 30 Μm (b, d, f).

Text-fig. 19. Scanning electron microscope (SEM; d, e) and synchrotron radiation X-ray tomographic microscopy (SRXTM; a–c) images of a seed of Kvacekispermum costatum sp. nov. (a–c) and Kvacekispermum sp. (d, e); Torres Vedras locality, Portugal. a, b) Isolated, abraded seeds showing densely spaced longitudinal ridges; c) Detail of abraded seed in (b) showing surface of endotesta with imprints of crystals; d) Isolated, abraded seed showing longitudinal ridges; e) Abraded seed surface showing surface of endotesta with imprints of crystals. Specimens, TV44-S174098 (holotype; a), TV38-S174099 (b, c), TV43-S174612 (d, e). Scale bars 300 Μm (a, b, d), 50 Μm (c, e). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 19. Scanning electron microscope (SEM; d, e) and synchrotron radiation X-ray tomographic microscopy (SRXTM; a–c) images of a seed of Kvacekispermum costatum sp. nov. (a–c) and Kvacekispermum sp. (d, e); Torres Vedras locality, Portugal. a, b) Isolated, abraded seeds showing densely spaced longitudinal ridges; c) Detail of abraded seed in (b) showing surface of endotesta with imprints of crystals; d) Isolated, abraded seed showing longitudinal ridges; e) Abraded seed surface showing surface of endotesta with imprints of crystals. Specimens, TV44-S174098 (holotype; a), TV38-S174099 (b, c), TV43-S174612 (d, e). Scale bars 300 Μm (a, b, d), 50 Μm (c, e).

Text-fig. 15. Scanning electron microscope (SEM) images of seeds of Pazliopsis reyi (a–c) and Anaspermum operculatum gen. et sp. nov. (d–j). a) Seed in lateral view showing poorly preserved remains of fruit wall and rough surface of the exotesta; b) Section through the wall of a broken seed showing palisade cells of the exotesta; c) Seed surface showing sunken very deeply sinuous outlines of the exotesta cells; d–f, h, i) Seeds in lateral view (holotype figured in d) showing the pointed micropylar-hilar area, the rounded chalazal end, the smooth surface of the exotesta composed of cells with sinuous cell outlines, and the distinct course of the raphe; g) Surface of the exotesta showing the distinctive cell outlines formed by the sinuous anticlinal walls of the palisade cells; j) Apical view of seed showing the prominent operculum. Specimens, TV43-S171534 (a), TV44-S136683 (b), TV43-S136745 (c), TV43-S136740 (holotype; d), TV43-S136746 (e), TV43-S136739 (f), TV43-S170083 (g), TV S136743 (h), TV43-S136742 (i), TV38-S174608 (j). Scale bars 300 Μm (a, b, d–f, h, i), 50 Μm (j), 30 Μm (c, g). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 15. Scanning electron microscope (SEM) images of seeds of Pazliopsis reyi (a–c) and Anaspermum operculatum gen. et sp. nov. (d–j). a) Seed in lateral view showing poorly preserved remains of fruit wall and rough surface of the exotesta; b) Section through the wall of a broken seed showing palisade cells of the exotesta; c) Seed surface showing sunken very deeply sinuous outlines of the exotesta cells; d–f, h, i) Seeds in lateral view (holotype figured in d) showing the pointed micropylar-hilar area, the rounded chalazal end, the smooth surface of the exotesta composed of cells with sinuous cell outlines, and the distinct course of the raphe; g) Surface of the exotesta showing the distinctive cell outlines formed by the sinuous anticlinal walls of the palisade cells; j) Apical view of seed showing the prominent operculum. Specimens, TV43-S171534 (a), TV44-S136683 (b), TV43-S136745 (c), TV43-S136740 (holotype; d), TV43-S136746 (e), TV43-S136739 (f), TV43-S170083 (g), TV S136743 (h), TV43-S136742 (i), TV38-S174608 (j). Scale bars 300 Μm (a, b, d–f, h, i), 50 Μm (j), 30 Μm (c, g).

Text-fig. 11. Scanning electron microscope (SEM) images of seeds assigned to the BEG group (a–d) and associated pollen grains (e–i); Torres Vedras locality, Portugal. a) Seed of Tomcatia taylorii showing the four horns formed by extensions of the envelope and the central projection of the envelope that surrounds to the micropylar tube; b, c) Seeds of Quadrispermum parvum in lateral (b) and apical (c) views showing the transverse ribs and the central projection of the envelope that surrounds the micropylar tube; d–f) Seeds of Ephedrispermum lusitanicum showing the four-angled seed envelope (d), the micropylar tube surrounded by the tissues of the integument (e), and ephedroid pollen grains on the seed surface (f); g) Apex of seed of Quadrispermum parvum showing simple in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 11. Scanning electron microscope (SEM) images of seeds assigned to the BEG group (a–d) and associated pollen grains (e–i); Torres Vedras locality, Portugal. a) Seed of Tomcatia taylorii showing the four horns formed by extensions of the envelope and the central projection of the envelope that surrounds to the micropylar tube; b, c) Seeds of Quadrispermum parvum in lateral (b) and apical (c) views showing the transverse ribs and the central projection of the envelope that surrounds the micropylar tube; d–f) Seeds of Ephedrispermum lusitanicum showing the four-angled seed envelope (d), the micropylar tube surrounded by the tissues of the integument (e), and ephedroid pollen grains on the seed surface (f); g) Apex of seed of Quadrispermum parvum showing simple

Text-fig. 14. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of seeds of Gastonispermum antiquum sp. nov. (a–e) and possible fruits (f–h); Torres Vedras locality, Portugal. a–c) Lateral view of seeds (holotype figured in b) showing the hilar-micropylar area and the course of the raphe; note the close association of hilum (arrowhead) and micropyle; d) Detail of seed wall showing the undulating surface of the exotesta composed of cells with sinuous cell outlines; e) Longitudinal section (orthoslice xy0914) through the hilar-micropylar area showing the hilum (arrowhead), the thickening of the endotesta around the micropyle and the course of the raphe within the exotesta (arrow); f–h) Fruits containing in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 14. Scanning electron microscope (SEM) and synchrotron radiation X-ray tomographic microscopy (SRXTM) images of seeds of Gastonispermum antiquum sp. nov. (a–e) and possible fruits (f–h); Torres Vedras locality, Portugal. a–c) Lateral view of seeds (holotype figured in b) showing the hilar-micropylar area and the course of the raphe; note the close association of hilum (arrowhead) and micropyle; d) Detail of seed wall showing the undulating surface of the exotesta composed of cells with sinuous cell outlines; e) Longitudinal section (orthoslice xy0914) through the hilar-micropylar area showing the hilum (arrowhead), the thickening of the endotesta around the micropyle and the course of the raphe within the exotesta (arrow); f–h) Fruits containing

Text-fig. 10. Scanning electron microscope (SEM) images of conifer seeds (a, b) and pollen (c) and monoporate pollen of unknown affinity (d–j); Torres Vedras locality, Portugal. a, b) Unnamed conifer seeds (conifer seed sp. 1); c) Clump of bisaccate pollen grains; d) Fragment with microsporangia that yielded the pollen in (e–j); e, f, h) Monoporate pollen grains folded in various ways, exposing the tiny pore (e, h, arrowheads) or resembling a monocolpate grain (f); g) Detail of pollen grain showing pore (arrowhead) and finely rugulate exine surface that reflects the reticulate infratectal layer beneath the thin tectum; i) Detail of pore showing very slightly thickened margin; j) Spherical orbicules on the surface of two grains. Specimens, TV44-S174594 (a), TV44-S174595 (b), TV44-S174573 (c), TV44-S137904 (d–j). Scale bars 1 mm (a, b), 300 Μm (d), 100 Μm (c), 6 Μm (e, f, h), 3 Μm (g, j), 1.5 Μm (i). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community

Text-fig. 10. Scanning electron microscope (SEM) images of conifer seeds (a, b) and pollen (c) and monoporate pollen of unknown affinity (d–j); Torres Vedras locality, Portugal. a, b) Unnamed conifer seeds (conifer seed sp. 1); c) Clump of bisaccate pollen grains; d) Fragment with microsporangia that yielded the pollen in (e–j); e, f, h) Monoporate pollen grains folded in various ways, exposing the tiny pore (e, h, arrowheads) or resembling a monocolpate grain (f); g) Detail of pollen grain showing pore (arrowhead) and finely rugulate exine surface that reflects the reticulate infratectal layer beneath the thin tectum; i) Detail of pore showing very slightly thickened margin; j) Spherical orbicules on the surface of two grains. Specimens, TV44-S174594 (a), TV44-S174595 (b), TV44-S174573 (c), TV44-S137904 (d–j). Scale bars 1 mm (a, b), 300 Μm (d), 100 Μm (c), 6 Μm (e, f, h), 3 Μm (g, j), 1.5 Μm (i).

Text-fig. 1. Stutzeliastrobus bohemicus (BAYER) J.KVAČEK, No. NM-F 2746, Harcov, lectotype. a – surface view of ovuliferous cone photograph, scale bar 10 mm, b – microCT isosurface of ovuliferous cone, scale bar 10 mm, c – microCT longitudinal section of ovuliferous cone with segmented seeds, scale bar 10 mm, d – microCT longitudinal section of ovuliferous cone in yellow, seeds in red, e – 3D visualised bract-scale complex bearing three seeds, adaxial view, scale bar 2.5mm, f – 3D visualised bract-scale complex bearing two seeds, lateral view (incomplete reconstruction of the scale visualises the front seed), scale bar 2.5mm. in Stutzeliastrobus Bohemicus Comb. Nov. - Basal Cupressaceae Conifer From The Cenomanian Of The Bohemian Cretaceous Basin, Central Europe

Text-fig. 1. Stutzeliastrobus bohemicus (BAYER) J.KVAČEK, No. NM-F 2746, Harcov, lectotype. a – surface view of ovuliferous cone photograph, scale bar 10 mm, b – microCT isosurface of ovuliferous cone, scale bar 10 mm, c – microCT longitudinal section of ovuliferous cone with segmented seeds, scale bar 10 mm, d – microCT longitudinal section of ovuliferous cone in yellow, seeds in red, e – 3D visualised bract-scale complex bearing three seeds, adaxial view, scale bar 2.5mm, f – 3D visualised bract-scale complex bearing two seeds, lateral view (incomplete reconstruction of the scale visualises the front seed), scale bar 2.5mm.

Text-fig. 3. Stutzeliastrobus bohemicus (BAYER) J.KVAČEK, Harcov. a – surface view of bract-scale complex (arrow) and probably aborted seed, No. NM-F 2746 (lectotype), scale bar 1 mm, b – seed with a wing (arrow) NM-F 872, scale bar 1 mm, c – microCT perpendicular section of bract scale complex showing two seeds with wings (arrows), No. NM-F 2746 (lectotype), scale bar 1 mm, d – microCT longitudinal section of bract scale complex showing three seeds, No. NM-F 2746 (lectotype), scale bar 1 mm, e – microCT longitudinal section of bract scale complex with one seed reconstructed showing micropyle (arrow), No. NM-F 2746 (lectotype), scale bar 1 mm, f – isolated seed with a fragment of wing (arrow), No. NM-F 2746 (lectotype), scale bar 1 mm. in Stutzeliastrobus Bohemicus Comb. Nov. - Basal Cupressaceae Conifer From The Cenomanian Of The Bohemian Cretaceous Basin, Central Europe

Text-fig. 3. Stutzeliastrobus bohemicus (BAYER) J.KVAČEK, Harcov. a – surface view of bract-scale complex (arrow) and probably aborted seed, No. NM-F 2746 (lectotype), scale bar 1 mm, b – seed with a wing (arrow) NM-F 872, scale bar 1 mm, c – microCT perpendicular section of bract scale complex showing two seeds with wings (arrows), No. NM-F 2746 (lectotype), scale bar 1 mm, d – microCT longitudinal section of bract scale complex showing three seeds, No. NM-F 2746 (lectotype), scale bar 1 mm, e – microCT longitudinal section of bract scale complex with one seed reconstructed showing micropyle (arrow), No. NM-F 2746 (lectotype), scale bar 1 mm, f – isolated seed with a fragment of wing (arrow), No. NM-F 2746 (lectotype), scale bar 1 mm.

Text-fig. 5. Reyispermum parvum gen. et sp. nov. seeds from the Early Cretaceous Vale de Água locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings). a) Holotype (S174178; Vale de Agua sample 141) in lateral view showing shape and cell pattern; remains of mounting media (¤). b) Cut volume rendering of seed (cut at yz0553) showing the slightly raised tissue immediately adjacent to the lower edge of the hilum (arrow head) and palisade-shaped cells of exotesta. c) Apical view of seed showing hilar depression (hi), position of micropylar slit (mi) and the slightly raised raphal ridge (ra). d) Seed in lateral view showing raised tissue immediately adjacent to the lower edge of the hilum (arrow head) (S174495, Vale de Água sample 300). e) Cut volume rendering (cut at yz0500) of the seed in (5d) showing the raised tissue (arrow head) immediately adjacent to the lower edge of the hilum and sclerenchyma cells of exotesta. f) Detail of seed in (5d) showing micropylar slit (mi), hilum (hi) and raised tissue (arrow head) immediately adjacent to the lower edge of the hilum. g, h) Seed in lateral view (g) and view towards raphe (h) showing seed shape, the raised tissue below hilum (arrow head) and the raphal ridge (ra); note pointed micropylar area (S174179, Vale de Água sample 141). i) Seed surface of seed in (5d) showing the raised outlines of the undulate anticlinal walls of the exotestal cells. Scale bars = 250 µm (a–e, g, h); 125 µm (f, i). in Extinct Taxa Of Exotestal Seeds Close To Austrobaileyales And Nymphaeales From The Early Cretaceous Of Portugal

Text-fig. 5. Reyispermum parvum gen. et sp. nov. seeds from the Early Cretaceous Vale de Água locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings). a) Holotype (S174178; Vale de Agua sample 141) in lateral view showing shape and cell pattern; remains of mounting media (¤). b) Cut volume rendering of seed (cut at yz0553) showing the slightly raised tissue immediately adjacent to the lower edge of the hilum (arrow head) and palisade-shaped cells of exotesta. c) Apical view of seed showing hilar depression (hi), position of micropylar slit (mi) and the slightly raised raphal ridge (ra). d) Seed in lateral view showing raised tissue immediately adjacent to the lower edge of the hilum (arrow head) (S174495, Vale de Água sample 300). e) Cut volume rendering (cut at yz0500) of the seed in (5d) showing the raised tissue (arrow head) immediately adjacent to the lower edge of the hilum and sclerenchyma cells of exotesta. f) Detail of seed in (5d) showing micropylar slit (mi), hilum (hi) and raised tissue (arrow head) immediately adjacent to the lower edge of the hilum. g, h) Seed in lateral view (g) and view towards raphe (h) showing seed shape, the raised tissue below hilum (arrow head) and the raphal ridge (ra); note pointed micropylar area (S174179, Vale de Água sample 141). i) Seed surface of seed in (5d) showing the raised outlines of the undulate anticlinal walls of the exotestal cells. Scale bars = 250 µm (a–e, g, h); 125 µm (f, i).

Text-fig. 3. Pazlia hilaris gen. et sp. nov. (a–e) from the Early Cretaceous Famalicão locality (sample 025), Portugal (holotype, S175096) and Pazliopsis reyi gen. et sp. nov. (f–i) from the Early Cretaceous Torres Vedras locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings a–f, i) and scanning electron microscopy (SEM, g, h). a, b) Seed in lateral (a) and oblique apical (b) views showing the truncate hilar-micropylar region; note prominent hilar scar (hi) and micropyle (mi) at the seed apex and the raphe (ra) seen as slightly raised ridge; remains of mounting media (¤). c) Cut volume rendering (cut at yz0647) showing course of raphe (ra), hilar scar (hi) and micropyle (mi); note the strongly radially elongated cells below the hilar scar. d) Seed in antiraphal view. e) Seed surface showing the raised undulate anticlinal walls of the exotestal cells. f) Seed enclosed in remains of thin-walled fruit (fr) (S174632, Torres Vedras sample 298). g) Holotype, seed enclosed in remains of fruit (fr); raphal view showing the faintly ribbed surface of the seed (S171534, Torres Vedras sample 043). h) Apical view of seed fragment showing hilar scar (hi), position of raphe (ra) and the ribbed seed surface (S136683, Torres Vedras sample 044). i) Seed surface showing the raised undulate anticlinal walls of the exotestal cells (S171534; Torres Vedras sample 043). Scale bars = 250 µm (a–d, f–h); 125 µm (e, i). in Extinct Taxa Of Exotestal Seeds Close To Austrobaileyales And Nymphaeales From The Early Cretaceous Of Portugal

Text-fig. 3. Pazlia hilaris gen. et sp. nov. (a–e) from the Early Cretaceous Famalicão locality (sample 025), Portugal (holotype, S175096) and Pazliopsis reyi gen. et sp. nov. (f–i) from the Early Cretaceous Torres Vedras locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings a–f, i) and scanning electron microscopy (SEM, g, h). a, b) Seed in lateral (a) and oblique apical (b) views showing the truncate hilar-micropylar region; note prominent hilar scar (hi) and micropyle (mi) at the seed apex and the raphe (ra) seen as slightly raised ridge; remains of mounting media (¤). c) Cut volume rendering (cut at yz0647) showing course of raphe (ra), hilar scar (hi) and micropyle (mi); note the strongly radially elongated cells below the hilar scar. d) Seed in antiraphal view. e) Seed surface showing the raised undulate anticlinal walls of the exotestal cells. f) Seed enclosed in remains of thin-walled fruit (fr) (S174632, Torres Vedras sample 298). g) Holotype, seed enclosed in remains of fruit (fr); raphal view showing the faintly ribbed surface of the seed (S171534, Torres Vedras sample 043). h) Apical view of seed fragment showing hilar scar (hi), position of raphe (ra) and the ribbed seed surface (S136683, Torres Vedras sample 044). i) Seed surface showing the raised undulate anticlinal walls of the exotestal cells (S171534; Torres Vedras sample 043). Scale bars = 250 µm (a–d, f–h); 125 µm (e, i).

Text-fig. 1. Gastonispermum portugallicum gen. et sp. nov. seeds from the Early Cretaceous Famalicão locality (sample 025), Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings). Note remains of mounting media on several seeds (¤). a) Seed in oblique view showing seed shape, the slightly raised raphal ridge and the position of hilum (hi) and micropyle (mi) on the raphal side of the seed (S170218). b, c) Seeds in lateral view (b, S170234; c, S175095). d–f) Holotype (S174820); seed in lateral view (d) and cut volume rendering (e, f) through the median plane of the seed showing palisade-shaped sclerenchyma cells of exotesta and remains of embryo (emb) and surrounding nutritive tissue (e, cut between yz0440-0530; f, cut between slices yz440-480). g) Hilum (hi) and micropyle (mi) of seed in (1a) showing the Y-shaped micropylar slit in the outer integument. h) Cut volume rendering through the median plane of the seed (cut at yz0492) showing seed coat mainly composed of palisade-shaped cells of the exotesta (S174435). i) Seed surface showing the raised outlines of the undulate anticlinal walls of the exotestal cells (S175045). Scale bars = 500 µm (a–e); 250 µm (g); 125 µm (f, i). in Extinct Taxa Of Exotestal Seeds Close To Austrobaileyales And Nymphaeales From The Early Cretaceous Of Portugal

Text-fig. 1. Gastonispermum portugallicum gen. et sp. nov. seeds from the Early Cretaceous Famalicão locality (sample 025), Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings). Note remains of mounting media on several seeds (¤). a) Seed in oblique view showing seed shape, the slightly raised raphal ridge and the position of hilum (hi) and micropyle (mi) on the raphal side of the seed (S170218). b, c) Seeds in lateral view (b, S170234; c, S175095). d–f) Holotype (S174820); seed in lateral view (d) and cut volume rendering (e, f) through the median plane of the seed showing palisade-shaped sclerenchyma cells of exotesta and remains of embryo (emb) and surrounding nutritive tissue (e, cut between yz0440-0530; f, cut between slices yz440-480). g) Hilum (hi) and micropyle (mi) of seed in (1a) showing the Y-shaped micropylar slit in the outer integument. h) Cut volume rendering through the median plane of the seed (cut at yz0492) showing seed coat mainly composed of palisade-shaped cells of the exotesta (S174435). i) Seed surface showing the raised outlines of the undulate anticlinal walls of the exotestal cells (S175045). Scale bars = 500 µm (a–e); 250 µm (g); 125 µm (f, i).