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Text-fig. 2. Plot of height versus height to width ratio of leaf scars of the studied species of Protopteris and Oncopteris. in Revision Of Protopteris And Oncopteris Tree Fern Stem Casts From The Late Cretaceous Of Central Europe
Text-fig. 2. Plot of height versus height to width ratio of leaf scars of the studied species of Protopteris and Oncopteris.
Text-fig. 1. Geological setting and location of the fossil sites mentioned in this paper. Grey area indicates Cretaceous Basins. in Revision Of Protopteris And Oncopteris Tree Fern Stem Casts From The Late Cretaceous Of Central Europe
Text-fig. 1. Geological setting and location of the fossil sites mentioned in this paper. Grey area indicates Cretaceous Basins.
FIG. 51 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 51. — Photographic documentation of field characters in Cyathea: A-E, habit; A'-E', trunk apex; A"-E", trunk surface at breast height; A-A", Cyathea madagascarica Bonap.; B-B", C. melanocaula Desv.; C-C", C. remotifolia Bonap.; D-D", C. concava Bonap.; E-E", C. dregei Kunze; A", persistent dead petiole bases removed from the trunk in the upper half of the picture; E, erect habit (above), creeping habit (below); E", branched creeping trunks. A-A", Janssen et al. 2829; B, E, uncollected; B', B", Janssen et al. 2608; C, C', Janssen et al. 2582; C", Janssen et al. 2432; D, D", Janssen et al. 2565; D', Janssen et al. 2579; E', Janssen et al. 2967; E", Janssen et al. 2779. Vouchers at P. Photos B', B", C', D' by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A-E, 1 m; A'-E', A"-E", 10 cm.
FIG. 45 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 45. — Distribution maps for Cyatheaceae (genus Cyathea) of Madagascar and the Comoros. Species are arranged in alphabetical order.
FIG. 42. — A-C in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 42. — A-C, Cyathea similis var. leptoderma Rakotondr. & Janssen; A, young sori with a membranous, globular indusium; B, mature sori with a rudimentary indusium forming a collar-like structure at the base of the receptacle; C, pinnule segments abaxially with a fragment of the costula; D-G, Cyathea similis var. montana Janssen & Rakotondr.; D, pinnule segments abaxially with a fragment of the costula, common form; E, pinnule segments abaxially with a fragment of the costula, large and strongly crenate form; F, pinnules adaxially with a fragment of the costa; G, scale from the base of the petiole. A-C, Rakotondrainibe 2703 (P); D, F, G, Guillaumet 4063 (P); E, Rakotondrainibe 3615 (P). Scale bars: A, B, 0.5 mm; C-E, G, 0.1 cm; F, 1 cm.
FIG. 53 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 53. — Photographic documentation of field characters in Cyathea: A-D, habit; A'-D', trunk apex; A"-D", trunk surface at breast height; A-A", Cyathea melleri (Baker) Domin; B-B", C. sechellarum Mett.; C-C", C. similis C.Chr.; D-D", C. valdesquamata Janssen & Rakotondr. A, A', B', B", uncollected; A", Janssen et al. 2569; B, Janssen et al. 3247; C, Janssen et al. 2402; C', Janssen et al. 2576; C", Janssen et al. 2393; D, D", Janssen et al. 2907; D', Janssen et al. 2904. Vouchers at P. Photos A'-A", C' by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A-D, 1 m; A'-D', A"-D", 10 cm.
FIG. 39 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 39. — Cyathea melleri (Baker) Domin: A, fertile pinnules abaxially with a fragment of the costa, sori only partly indicated; B, sterile pinnules abaxially with a fragment of the costa; C, pinnule segments abaxially with a fragment of the costula (at the junction with the costa), sporangia partly removed to reveal the hemitelioid indusia; D, habit; E, leaf scars and trunk surface; F, basal part of the leaf (from the petiole base up to the first pinna pair), basal part in lateral view, folded middle part in adaxial and upper part in dorsal view, note the presence of a pair of reduced pinnae at 5-10 cm above the base of the petiole; G, pinnule segments adaxially with a fragment of the costula (at the junction with the costa), form with narrow segments; H, scales from the base of the petiole (left: lateral view; right: dorsal view). A-C, E, F, H, Janssen et al. 2569 (P); D, uncollected, photograph at P; G, Rakotondrainibe et al. 5866 (P). Scale bars: A, B, 1 cm; C, H, 0.1 cm; D, 1 m; E, F, 5 cm; G, 0.5 cm.
FIG. 40 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 40. — Cyathea sechellarum Mett.: A, pinnules abaxially with a fragment of the costa, sori omitted from one pinnule; B, sori, note the cup-shaped indusium and scaly indument on the segment midvein; C, habit (left: adult plant; right: young plant); D, pinnule segments abaxially with a fragment of the costula; E, filiform scale from the abaxial face of a segment midvein; F, deltoid scale from the abaxial face of a segment midvein; G, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view, one half of the petiole pruned away by longitudinal section, note presence of a pair of reduced pinnae near the base of the petiole; H, scale from the base of the petiole; I, leaf scars and trunk surface, persistent dead petiole bases (above) removed from two scars (below). A, B, D, G, H, Janssen et al. 3226 (P); C (left), Janssen et al. 3247 (P), C (right), uncollected, photograph at P; E, F, Janssen et al. 3247 (P); I, Janssen et al. 3201 (P). Scale bars: A, G, 5 cm; B, 0.1 cm; C, 1 m; D, H, 0.5 cm; E, F, 0.5 mm; I, 1 cm.
FIG. 37 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 37. — Cyathea kirkii Hook.: A, pinnules abaxially with a fragment of the costa, sori omitted from one pinnule; B, pinnule segments abaxially from the basal part of a pinnule (at the junction of the costula with the costa), form with smaller segments; C, habit; D, pinnule segments abaxially from the basal part of a pinnule (at the junction of the costula with the costa), common form; E, basal part of the leaf (from the petiole base up to the first pinna pair), dorso-lateral view, ventral view in upper third, one half of the petiole pruned away by longitudinal section; F, scale from the base of the petiole (left: dorsal view; right: lateral view). A, D-F, Rakotondrainibe et al. 6729 (P); B, C, Labat et al. 3208 (P). Scale bars: A, E, 5 cm; B, D, F, 0.5 cm; C, 1 m.
FIG. 35 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 35. — Cyathea concava Bonap.: A, pinnule segments abaxially with a fragment of the costula (at the junction with the costa); B, fertile pinnules abaxially with a fragment of the costa, sori only partly indicated; C, sterile pinnule abaxially with a fragment of the costa; D, habit; E, scale from the base of the petiole; F, basal part of the leaf (from the petiole base up to the first pinna pair, one pinna pruned), lateral view; G, leaf scars and trunk surface. A, B, D, F, G, Janssen et al. 2565 (P); C, Janssen et al. 2564 (P); E, Janssen et al. 2579 (P). Scale bars: A, E, 0.5 cm; B, C, G, 1 cm; D, 1 m; F, 5 cm.
FIG. 47 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 47. — Photographic documentation of field characters in Cyathea: A-E, habit; A'-E', trunk apex; A"-E", trunk surface at breast height. A-A", Cyathea marattioides Willd. ex Kaulf.; B-B", C. perrieriana C.Chr.; C-C", C. auriculata Tardieu; D-D", C. decrescens Mett. ex Kuhn; E-E", C. ivohibensis (C.Chr.) Janssen & Rakotondr. A, Janssen et al. 2488; A', B', uncollected; A", Janssen et al. 2484; B, Janssen et al. 2504; B", Janssen et al. 2505; C-C", Janssen et al. 2508; D, Janssen et al. 2521; D', Janssen et al. 2562; D", Janssen et al. 2406; E, E', Janssen et al. 2790; E", Janssen et al. 2794. Vouchers at P. Photos B-D, A'-D', A"-D" by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A-E, 1 m; A'-E', A", B", D", E", 10 cm; C", 5 cm.
FIG. 34. — A-F, H-L in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 34. — A-F, H-L, Cyathea boivinii Mett. ex Kuhn var. hildebrandtii; A, pinnules abaxially with a fragment of the costa, common form, sori only partly indicated; B, pinnules abaxially with a fragment of the costa, form with broader pinnule segments, sori only partly indicated; C, pinnule segments abaxially with a fragment of the costula, form with broader pinnule segments; D, E, pinnule segments abaxially with a fragment of the costula, common forms; F, habit; H, sori, common form with long receptacles and cup-shaped indusia with entire rims; I, scales from the base of the petiole, form with narrow, blackish scales; J, scales from the base of the petiole, common form; K, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view, one half of the petiole pruned away by longitudinal section, note the presence of a pair of reduced pinnae at about 20 cm above the base of the petiole; L, leaf scars and trunk surface; G, C. boivinii var. parahildebrandtii Janssen & Rakotondr., mature sori with dehiscing indusia. A, D, H, J, K, Rakotondrainibe 1666 (P); B, C, Rakotondrainibe 4709 (P); E, I, Janssen et al. 2956 (P); F, Janssen et al. 2442 (P); G, Rakotondrainibe et al. 6730 (P); L, Janssen et al. 2464 (P). Scale bars: A, B, L, 1 cm; C-E, G, H, 0.1 cm; F, 1 m; I, J, 0.5 cm; K, 5 cm.
FIG. 52 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 52. — Photographic documentation of field characters in Cyathea: A-D, F, habit; A'-D', F', trunk apex; A"-D", F", trunk surface at breast height; A-A", Cyathea boivinii Mett. ex Kuhn var. boivinii; B-B", C. boivinii var. hildebrandtii (Kuhn) Janssen & Rakotondr.; C-C", C. boivinii var. andringitrensis Janssen & Rakotondr.; D, D', C. kirkii Hook.; F-F", C. lastii Baker; A", the petiole bases are sometimes completely decayed on older trunks (left), but persistent in most trunks observed (right); E, C. boivinii var. andringitrensis Janssen & Rakotondr., dense tomentum of intricate squamules on the petiole; A, A', A" (left), C, C', uncollected; A" (right), Janssen et al. 2604; B, Janssen et al. 2433; B', Janssen et al. 2442; B", Janssen et al. 2464; C", Janssen et al. 2594; D, D', Labat et al. 3208; E, Janssen et al. 2591; F-F", Janssen et al. 2431. Vouchers at P. Photos A', A" (left), B', B", C-C", E by G. Rouhan (MNHN); D, D' by J.-N. Labat (MNHN); all other photos by T. Janssen. Scale bars: A-D, F, 1 m; A'-D', F', A"-C", F", 10 cm; E, 1 cm.
FIG. 31 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 31. — Cyathea remotifolia Bonap.: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, pinnules abaxially with a fragment of the costa, sori only partly indicated; C, habit; D, sori on the abaxial face of a pinnule, the sorus in the centre with an irregularly dehisced indusium, all other sori with showing rudimentary indusia usually reduced to a collar-like structure at the base of the receptacle; E, basal part of the leaf (from the petiole base up to the fifth pinna pair); F, scales from the base of the petiole (left: a group of agglutinated scales; right: a single scale); G, leaf scars with prominent spines and smooth trunk surface. A, B, D, E, Janssen et al. 2432 (P); C, F, Janssen et al. 2582 (P); G, Janssen et al. 2443 (P). Scale bars: A, E, 5 cm; B, G, 1 cm; C, 1 m; D, 0.1 cm; F, 0.5 cm.
FIG. 46 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 46. — Distribution maps for Cyatheaceae (genus Cyathea) of Madagascar and the Comoros. Species are arranged in alphabetical order.
FIG. 30. — A-F in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 30. — A-F, Cyathea pseudobellisquamata Janssen & Rakotondr.; A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view; C, pinnules abaxially with a fragment of the costa; D, scale from the base of the petiole; E, trunk segment with leaf scars; F, part of a pinnule abaxially, note the close standing sori and small scales on the costula; G-J, C. simulans (Baker) Janssen & Rakotondr.; G, pinnae adaxially with a fragment of the rachis; H, scale from the petiole; I, pinnules abaxially with a fragment of the costa, sori omitted from one pinnule; J, one sorus with dense paraphyses much longer than the sporangia, indusium absent, one catenate paraphysis on the left. A-F, Rakotondrainibe 3059 (P); G-J, Perrier de la Bâthie 7444 (P). Scale bars: A, B, G, 5 cm; C, E, I, 1 cm; D, H, 0.5 cm; F, J, 0.1 cm.
FIG. 38 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 38. — Cyathea lastii Baker: A, pinnules adaxially with a fragment of the costa; B, pinnule segments abaxially with a fragment of the costula; C, part of a pinnule segment abaxially with sori, dorso-lateral view, note the cup-shaped indusium and the distance between adjacent sori; D, habit; E, leaf scars and trunk surface; F, scale from the base of the petiole; G, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view, one half of the petiole pruned away by longitudinal section. A-C, F, G, Janssen et al. 2430 (P); D, Janssen et al. 2431 (P); E, Janssen et al. 2381 (P). Scale bars: A, E, 1 cm; B, 0.5 cm; C, F, 0.1 cm; D, 1 m; G, 5 cm.
FIG. 29 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 29. — Cyathea orthogonalis Bonap.: A, pinnae abaxially with a fragment of the rachis; B, patent, petiolulate pinnae near the base of the lamina (arrangement reconstructed from the herbarium specimen); C, basal part of the leaf (from the petiole base up to the first two pinna pairs), lateral view, all pinnae folded to one side of the rachis; D, scale from the abaxial face of the lower part of the rachis, about 30 cm above the base of the petiole; E, scale from the lateral face of the lower part of the rachis, about 30 cm above the base of the petiole; F, scale from the abaxial face of the base of the petiole; G, pinnules abaxially with a fragment of the costa; H, sori; I, trunk with leaf scars. A-C, G-I, Rakotondrainibe et al. 3754 (P); D-F, DuPuy et al. M805 (P). Scale bars: A-C, 5 cm; D-F, H, 0.1 cm; G, I, 1 cm. A, B, G, I, drawings by B. Raufeisen.
FIG. 28. — A-F in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 28. — A-F, Cyathea melanocaula Desv.; A, basal part of the leaf (from the petiole base up to the first several pinna pairs), basal part in lateral view, upper part in adaxial view, folded part in abaxial view; B, leaf scars and trunk surface; C, habit; D, scales from the base of the petiole (left: dorsal view; right: lateral view); E, pinnae abaxially with a fragment of the rachis, sori omitted from one pinna; F, pinnules abaxially with a fragment of the costa; G-K, C. obtecta Rakotondr. & Janssen; G, pinnae abaxially with a fragment of the rachis, sori only partly indicated; H, pinnules abaxially with a fragment of the costa; I, detail from the base of a pinnule, note the obliquely cup-shaped indusia and dense scaly indument on the costa and costula; J, scale from the base of the petiole; K, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view, note the two pairs of aphlebioid pinnae near the base of the petiole. A-F, Janssen et al. 2608 (P); G-K, Rakotondrainibe 2524 (P). Scale bars: A, E, G, K, 5 cm; B, F, H, 1 cm; C, 1 m; D, 0.5 cm; I, J, 0.1 cm.
FIG. 27 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 27. — Cyathea madagascarica Bonap.: A, pinnules abaxially with a fragment of the costa; B, adaxial view of the rachis from the lower half of the leaf, note that the proximal pinnules overlap the sparsely scaly rachis; C, habit; D, scale from the abaxial face of the base of the rachis; E, basal part of the leaf (from the base of the petiole up to the first several pinna pairs), dorsal view; F, pinnae abaxially with a fragment of the rachis, sori only partly indicated; G, leaf scars and trunk surface. Janssen et al. 2829 (P). Scale bars: A, B, E, G, 1 cm; C, 1 m; D, 0.1 cm; F, 5 cm.
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