FIGURE 4. Uktena riparia n in Uktena riparia n. gen., n. sp. (Annelida, Clitellata, Lumbriculidae), a new spermatophore-producing oligochaete

FIGURE 4. Uktena riparia n. sp. from Flat Creek (the type locality), from sagittal sections (A–E and J–L), or dissections (F– I). A. Ectal part of spermathecal ampulla, with sperm lined up along epithelium. B. Ental part of spermathecal ampulla with weakly-stained (old?) sperm. C. Copulatory organ and gland in spermathecal bursa. D. Muscle layer of spermathecal bursa wall. E. Small glands and muscle layer forming the wall of male bursa. F. Gland opening on papilla within male bursa. G. Spermatophore forming in atrial duct, with darkly-staining sperm (composite photo). H. Atrial ampulla and duct with coiled sperm bundles inside. I. Spermatophore protruding from male pore. J–L. Atrial ampullae; (K) showing detail of atrial wall, with prostate glands penetrating muscle layer (arrows); (L) showing elongate prostate glands with secretions.