FIGURE 3. Paranadrilus descolei Gavrilov, 1955. A–G,I,J in On spermatophore-producing aquatic microdrile oligochaetes (Annelida: Clitellata)

FIGURE 3. Paranadrilus descolei Gavrilov, 1955. A–G,I,J from Marchese's collection, H,K,L from Gavrilov's collection. A. Anterior part of the body. B. Ventral gland in the posterior part of segment V. C. Spermatophores within a lateral pouch in the anterior part of segment XII. D. Detail of the attachment of the spermatophores within the pouch. E. Transverse section of the empty spermatophore pouch. F. Sperm funnel and male pore slit. G. Bulbous male sac containing the copulatory organ, and empty spermatophore pouch behind septum 11/12. H. Scarcely developed male sac containing a long copulatory organ covered by a chitinous layer in a dissected specimen from Gavrilov's type collection. I. Male sac with copulatory organ and copulatory bursa. J. Spermatophore in the ental part of the copulatory bursa. K,L. Segment XI of the lectotype of P. descolei, showing the large ovary, the bulbous male sac filled by glandular secretions and spermatozoids, and the copulatory bursa with folded walls. A,E,G,I–L, histological sections. Abbreviations: at attachment site, cb copulatory bursa, co copulatory organ, ed ejaculatory duct, m muscular layer, mp male pore, ms male sac, o ovary, ph pharynx, sf sperm funnel, sp spermatophore pouch, sph spermatophore, vd vas deferens.