FIGURE 109. A in Three new species of Olivierus (Scorpiones: Buthidae) from Kazakhstan and Uzbekistan

FIGURE 109. A time-calibrated phylogeny for Olivierus species in Central Asia: Olivierus mikhailovi sp. n., O. tarabaevi sp. n., O. voldemari sp. n., and O. gorelovi (adapted from Graham et al., 2019, fig. 4). The tree is based on two mitochondrial (COI & 16S) and one nuclear (ITS-2) markers. Horizontal bars represent highest posterior densities (95%) around mean date estimates. Nodes with black dots were supported with high posterior support (>95). Vertical bars represent different species based on the most conservative consensus of three species delimitation approaches (PTP, GMYC, & ABGD). Abbreviations: KZ, Kazakhstan; TM, Turkmenistan; UZ, Uzbekistan. See Graham et al. (2019) for additional information.