Figure 5. Legs. A–E in The evolution of Bolbites onitoides (Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem

Figure 5. Legs. A–E, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. C–E, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina.