Figure 4 in The evolution of Bolbites onitoides (Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem

Figure 4. Meso- and metaventrite in Bolbites and Gromphadina. A, Bolbites onitoides. B, Gromphas inermis. C, D, Oruscatus davus: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most Bolbites and and the other Phanaeina whose mesoventral ridge is T-shaped. In Oruscatus, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between Bolbites and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity.