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FIGURE 26. Alyonushka echinata n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 26. Alyonushka echinata n. gen., n. sp. A–C, holotype, ZIRAS 1/50712, rotational views of fertile colony; D–F, paratype, ZIRAS 2/50713, rotational views of fertile colony. Arrows indicate the ooeciostome in all photos. Scale bars: 250 µm.

opennotspecifiedSep 2018View details →
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FIGURE 47. Haywardozoon pacificum n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 47. Haywardozoon pacificum n. sp. Specimens stained in Rose Bengal. A, specimen GLD4–11, Stn 214, zooids on a serpulid tube; B–G, holotype, ZIRAS 1/50726, different views of zooids on a serpulid tube; H–J, specimen GLD4–12, Stn 245, respectively showing very lightly stained zooids on serpulid tube and two zooids seen in transmitted light. Abbreviations: bb, brown body; d, diaphragm; dmd, dilator muscles of diaphragm; g, gut; olf, operculum-like flap; rmtc, retractor muscles of tentacle crown; t, tentacle cluster; v, vestibule. Scale bars: A–H, 500 µm; I, J, 100 µm.

opennotspecifiedSep 2018View details →
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FIGURE 23. Alyonushka filia n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 23. Alyonushka filia n. gen., n. sp. Progressive stages of development of ancestrulate and slightly older colonies. A, B, specimen GLD4–09, Stn 173, at four-zooid stage, with alveoli; C, D, specimen YMG4–14, Stn 363, four-zooid colony with one peristome occluded and spikes on trabeculae; E, F, specimen GLD4–11, Stn 224, also at four-zooid stage but with broader base; G, H, specimen YMG4–14, Stn 365 with more-convex colony center; I, J, specimen YMG4–14, Stn 351, nearsymmetrical four-zooid colony; K, L, specimen GLD4–09, Stn 180, colony with disproportionately thickened base and irregular column margin, suggesting possible regeneration after damage; M, N, specimen GLD4–12, Stn 235, an asymmetrical five-zooid colony; O, P, specimen YMG4–13, Stn 294, with unusually thick basal support. Scale bars: 200 µm.

opennotspecifiedSep 2018View details →
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FIGURE 25. Alyonushka hystricosa n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 25. Alyonushka hystricosa n. gen., n. sp. Holotype, ZIRAS 1/50711. A, ooeciostome; B, calyx margin with autozooidal peristomes, ooeciopore near middle right; C, close-up of peristomes and surface sculpture, ooeciopore at lower right; D, colony base; notice minutely prickled surface; E, oblique view of calyx, ooeciostome to right of center; F, symmetrical alveolus opening between a pair of peristomes. Scale bars: 100 µm.

opennotspecifiedSep 2018View details →
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FIGURE 45 in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 45.?Anyutidae sp. incertae sedis. A–D, rotational views of colony ZIRAS 1/50725; note the highly alveolate colony surface and fascicles of variable size. Scale bars: 500 µm.

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FIGURE 52. Alyonushka filia n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 52. Alyonushka filia n. gen., n. sp. Micro-CT scans of paratype, NIWA 127724, as back-face isosurface renders showing colony interiors. A, entire colony in profile; note the density of kenozooidal chambers in the column; B, close-up of distal part of column showing diminished numbers of kenozooids where it broadens into the calyx; C, optical section through gonozooid and ooeciostome, with arrowhead indicating a ledge at inner opening of ooeciostome; D, optical section through base of column showing protoecial dome with ancestrular peristome emerging from its apex (purple) and first daughter zooid (red) also originating from protoecium (its communication pore outside plane of section); note multiple layers of kenozooids; E, three successive sections through ancestrular region, moving upwards from left to right, respectively showing protoecium (red) and first daughter zooid (yellow), narrowing to ancestrular peristome and broadening peristome of daughter zooid. Scale bars: A, B, 250 µm; C–E, 100 µm.

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FIGURE 21. Alyonushka filia n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 21. Alyonushka filia n. gen., n. sp. A, holotype, ZIRAS 1/50710, profile of colony with paired gonozooid; B, paratype, NIWA 127724, profile of colony with single gonozooid; C–E, specimen YMG4–13, Stn 299, bilaterally compressed, showing developing gonozooid with continuous floor. Scale bars: A, B, 500 µm; C–E, 250 µm.

opennotspecifiedSep 2018View details →
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FIGURE 22. Alyonushka filia n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 22. Alyonushka filia n. gen., n. sp. A–E, G–L, holotype, ZIRAS 1/50710; E, F, paratype, NIWA 127724. A, frontal (apical) view of colony showing two gonozooids separated by a narrow cleft, each chamber with an ooeciopore (arrows); B, oblique view of A from 8 o'clock perspective, showing two gonozooid mounds separated by cleft (concealed in furrow), each mound with complex spiky alveolate surface; C, same, from different angle, cleft at middle left; D, autozooidal peristomes with linear ridges and barbs, and pores; E, skeletal microstructure of gonozooid surface; F, I, incompletely developed autozooidal peristomes with abundant internal spinules; G, close-up of part of cleft between adjacent gonozooids; H, base of colony with hummocks indicating underlying alveolar/kenozooid-like chambers; note densely prickled surface; J, L, ooeciostomes of the two gonozooids shown in A; K, apical view of mature autozooidal peristome. Scale bars: A, 500 µm; B–D, H, 250 µm; F, G, I, J, L, 100 µm; E, K, 50 µm.

opennotspecifiedSep 2018View details →
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FIGURE 38 in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 38. Genus et species indet. Progressive stages of development of ancestrular and young colonies. A–D, specimen GLD4–09, Stn 183, six-zooid stage, with peristomes and alveoli bounded by extrazooidal space(s); later stages shown in E–H (specimen YMG4–06, Stn 80), I–L (specimen GLD4–09, Stn 192) and M–P (specimen YMG18–01, Stn 38). Scale bars: 200 µm.

opennotspecifiedSep 2018View details →
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FIGURE 24. Alyonushka hystricosa n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 24. Alyonushka hystricosa n. gen., n. sp. A, B, holotype, ZIRAS 1/50711, apical and oblique views of capitulum with gonozooid; C, D, same specimen showing entire colony. Arrows indicate the same ooeciostome in each photo. Scale bars: 250 µm.

opennotspecifiedSep 2018View details →
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FIGURE 34. Calyssopora clarionensis n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG4–13, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD4–11, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: A–F, 200 µm; G–I, 100 µm; J–P, 50 µm.

opennotspecifiedSep 2018View details →
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FIGURE 20. Saccocamera ampulla n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 20. Saccocamera ampulla n. gen., n. sp. Holotype, ZIRAS 1/50709. A, Frontal (apical) view of gonozooid, with peristome at 4 o'clock; B, lateral view of gonozooid, peristome directed to right; note the numerous alveolar openings; C, D, gonozooid showing lageniform ooeciostome; E, ooeciopore; F–J, profiles of autozooidal peristomes showing surface texture and spinose processes; K, autozooidal aperture. Scale bars: A, B, 200 µm; C, D, F–J, 100 µm; E, K, 50 µm.

opennotspecifiedSep 2018View details →
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FIGURE 41. Anyuta anastema n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm.

opennotspecifiedSep 2018View details →
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FIGURE 54. Rallocytus ridiculus n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 54. Rallocytus ridiculus n. gen., n. sp. Micro-CT scans of paratype 2, NIWA 127726, as surface (A) and back-face isosurface renders showing colony interiors. A, surface and interior views of seven-zooid colony with dimorphic ancestrular zooid (daz) colored purple in the right-hand image; note small cluster of flattened kenozooidal chambers surrounding protoecium but absent from sides of column; B, lateral view, with entire dimorphic ancestrular zooid seen in left half; C, apical view of calyx, showing flattened kenozooidal/alveolar chambers filling center, dimorphic peristome at right indicated by arrowhead. Scale bars: 100 µm.

opennotspecifiedSep 2018View details →
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FIGURE 32. Calyssopora vasiformis n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 32. Calyssopora vasiformis n. gen., n. sp. Stages of colony development. A–D, specimen GLD4–11, Stn 228, at apparent four-zooid stage with large alveoli; E–H, specimen GLD4–11, Stn 201, at a later stage, with more-developed calyx center. Scale bars: 200 µm.

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FIGURE 44. Rallocytus ridiculus n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 44. Rallocytus ridiculus n. gen., n. sp. Progressive stages of development of ancestrulate colonies. A–D, specimen YMG4–14, Stn 340, at three-zooid stage, with no distinction among peristomes; E–H, specimen GLD4–09, Stn 183, with four functional zooids, one dimorphic; I–L, specimen GLD4–08, Stn 159, with six functional zooids, one dimorphic; M–P, specimen GLD4–11, Stn 206, five functional zooids, none dimorphic. Scale bars: A, E, 150 µm; B–D, F–P, 200 µm.

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FIGURE 19. Saccocamera ampulla n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 19. Saccocamera ampulla n. gen., n. sp. A–D, four profiles of fertile holotype colony, ZIRAS 1/50709, YMG4–14, Stn 359. Scale bars: 250 µm.

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FIGURE 51. Orectopora flabellum n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 51. Orectopora flabellum n. gen., n. sp. Micro-CT scans of paratype, NIWA 127723, as back-face isosurface renders showing colony interiors. A, colony curvature in lateral view showing part of abfrontal (left) and frontal (right) surfaces; note large cluster of kenozooidal chambers at lower left, becoming longer distally and near autozooidal tubes; B, like A, but from opposite side, showing more of abfrontal surface; note pseudopore density; C, en face view of colony frontal surface, with only the part in the square shown as backface isosurface render; note the kenozooids in the interior angle between two-right-hand fascicles (lobes); D, E, abfrontal side of colony, with that in D showing differentiating tip of middle lobe; note elongate kenozooids on either side of or abfrontal to autozooidal tubes; F, close-up of part of A showing variability of kenozooidal chambers. Scale bars: 100 µm.

opennotspecifiedSep 2018View details →
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FIGURE 17. Frontohornera frontalis n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm.

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FIGURE 18. Frontohornera frontalis n. gen., n in Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining

FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. A–C, specimen GLD4–11, Stn 215, from different angles; D–F, specimen YMG4–14, Stn 334, from different angles; G–I, specimen GLD4–11, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: A–H, 500 µm; I, 250 µm.

opennotspecifiedSep 2018View details →

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International Brain Laboratory public data

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Last verified 2026-04-29Open record