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FIG. 26 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 26. — Cyathea longispina Janssen & Rakotondr.: A, pinnae with a fragment of the rachis (above: adaxially; below: abaxially, sori only partly indicated); B, habit; C, proximal part of a pinnule abaxially (at the junction with the costa), note the scaly indument on the costa and costula; D, pinnules abaxially with a fragment of the costa; E, scale from the abaxial face of the costa, note the long apical spine; F, pinnule apex adaxially with straight multicellular hairs on the costula and crispate catenate hairs on the veins, both types of hairs two-fold magnified on the left; G, scale from the base of the petiole; H, scale from the upper half of the petiole; I, basal part of the leaf (from the petiole base up to the second pinna pair), dorsal view, pinnae pruned away on one side; J, leaf scars and trunk surface. A, C-J, Janssen et al. 2916 (P); B, Janssen et al. 2908 (P). Scale bars: A, I, 5 cm; B, 1 m; C, E, F, 0.1 cm; D, J, 1 cm; G, H, 0.5 cm.
FIG. 24 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 24. — Cyathea emilei var. dauphinensis Janssen & Rakotondr.: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, pinnule abaxially with a fragment of the costa, crenate form; C, pinnules abaxially with a fragment of the rachis, common form; D, basal part of the leaf (from the petiole base up to the second pinna pair), lateral view; E, scale from the base of the petiole; F, group of scales from the base of the petiole, their upper half stramineous and membranous; G, scale from the lateral-adaxial face of the middle part of the petiole; H, leaf scars and trunk surface. A, C-E, G, Janssen et al. 2617 (P); B, F, H, Janssen et al. 2612 (P). Scale bars: A, D, 5 cm; B, C, H, 1 cm; E-G, 0.5 cm.
FIG. 33. — A-G in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 33. — A-G, Cyathea boivinii Mett. ex Kuhn var. andringitrensis Janssen & Rakotondr.; A, pinnule segments abaxially with a fragment of the costula; B, scale from the base of the petiole; C, basal part of the leaf (from the petiole base up to the second pinna pair), lateral view, the scales are caducous; D, leaf scars and trunk surface; E, habit of a crown with simultaneously unrolling young leaves; F, pinnules abaxially with a fragment of the costa, sori only partly indicated; G, habit, note the heap of dead leaves at the base of the trunk from the last simultaneous shedding; H-J, Cyathea boivinii var. bevolo Janssen & Rakotondr.; H, basal part of the leaf (petiole base up to the first pinna pair), lateral view; I, pinnule segments abaxially with a fragment of the costula; J, pinnules abaxially with a fragment of the costa, sori only partly indicated. A-C, F, Janssen et al. 2600 (P); D, Janssen et al. 2594 (P); E, G, uncollected, photographs at P; H-J, Perrier de la Bâthie 13508 (P). Scale bars: A, I, 0.1 cm; B, 0.5 cm; C, H, 5 cm; D, F, J, 1 cm; E, G, 1 m.
FIG. 25 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 25. — Cyathea longipinnata Bonap.: A, pinnae abaxially with a fragment of the rachis; B, habit; C, pinnules abaxially with a fragment of the costa, common form; D, pinnules abaxially with a fragment of the rachis, small form with rounded apex; E, adaxial view of the rachis from the lower half of the leaf, note that the proximal pinnules overlap the rachis; F, scale from the lateral-adaxial face of the lower half of the rachis; G, scale from the abaxial face of the base of the rachis; H, leaf scars and trunk surface; I, basal part of the leaf (from the base of the petiole up to the first several pinna pairs), dorsal view. A-C, E, I, Janssen et al. 2557 (P); D, F-H, Janssen et al. 2507 (P). Scale bars: A, I, 5 cm; B, 1 m; C-E, H, 1 cm; F, G, 0.1 cm. A, C, I, drawings by B. Raufeisen.
FIG. 23 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 23. — Cyathea emilei Janssen & Rakotondr.: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, pinnules abaxially with a fragment of the costa, common form; C, pinnules abaxially with a fragment of the costa, form with small pinnules with rounded apices; D, habit; E, basal part of the leaf (from the petiole base up to the second pinna pair), lateral view in lower, dorsal view in upper half; F, pinnules abaxially with a fragment of the costa, form with big pinnules with acute apices, sori omitted from two pinnules; G, proximal part of a pinnule abaxially (at the junction with the costa), note the persistent, coriaceous indusia; H, group of agglutinated scales from the base of the petiole; I, scale from the base of the petiole (left: common form, light brown and caducous; right: dark brown and persistent form); J, leaf scars and trunk surface. A, B, D, E, G-I (left), Janssen et al. 2580 (P); C, J, Janssen et al. 2567 (P); F, I (right), Janssen et al. 2959 (P). Scale bars: A, E, 5 cm; B, C, F, J, 1 cm; D, 1 m; G, 0.1 cm; H, I, 0.5 cm.
FIG. 22. — A-D in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 22. — A-D, Cyathea bellisquamata var. australis Janssen & Rakotondr.; A, pinnules abaxially with a fragment of the costa; B, scale from the base of the petiole; C, segment from the basal part of the petiole with scales; D, proximal part of a pinnule abaxially (at the junction with the costa); E-H, C. bellisquamata var. basilobata C.Chr.; E, pinnules abaxially with a fragment of the costa; F, scale from the base of the petiole; G, proximal part of a pinnule abaxially (at the junction with the costa); H, segment from the basal part of the petiole with scales. A-D, Rakotondrainibe et al. 4331 (P); E-H, Janssen et al. 2937 (P). Scale bars: A, C, E, H, 1 cm; B, D, F, G, 0.1 cm.
FIG. 21 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 21. — Cyathea bellisquamata Bonap. var. bellisquamata: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, habit; C, pinnules abaxially with a fragment of the costa, common form; D, pinnules abaxially with a fragment of the costa, small form; E, proximal part of a pinnule abaxially (at the junction with the costa), note the dark, coriaceous indusia; F, scale from the base of the petiole; G, basal part of the leaf (from the petiole base up to the first nine pinna pairs), lateral view, one half of the petiole pruned away by longitudinal section; H, leaf scars and trunk surface. A, C, E-G, Janssen et al. 2482 (P); B, H, Janssen et al. 2522 (P); D, Humbert 22996 (P). Scale bars: A, G, 5 cm; B, 1 m; C, D, H, 1 cm; E, F, 0.1 cm.
FIG. 50 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 50.— Photographic documentation of field characters in Cyathea: A-E, habit;A'-E', trunk apex; A"-E", trunk surface at breast height. A-A", Cyathea bellisquamata Bonap.; B-B", C. emilei Janssen & Rakotondr. avr. emilei; C-C", C. emilei var. dauphinensis Janssen & Rakotondr.; D-D", C. longipinnata Bonap.; E-E", C. longispina Janssen & Rakotondr. A-A", Janssen et al. 2522; B, B', Janssen et al. 2580; B", Janssen et al. 2567; C, C", Janssen et al. 2612; C', uncollected; D-D", Janssen et al. 2507; E, Janssen et al. 2908; E', Janssen et al. 2915; E", Janssen et al. 2916. Vouchers at P. Photos A-D, A'-D', A"-D" by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A-E, 1 m; A'-E', C"-E", 10 cm; A", B", 5 cm.
FIG. 43 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 43. — Cyathea valdesquamata Janssen & Rakotondr.: A, leaf scars and trunk surface; B, pinnule segments abaxially with a fragment of the costula (at the junction with the costa), note the dense scaly indument; C, habit; D, rachis fragment with the basal part of two pinnae, abaxial view, sori only partly indicated, note the close standing pinnules and the proximal pinnules overlapping the rachis; E, scale from the abaxial face of a costula; F, basal part of the leaf (from the petiole base up to the first seven pinna pairs), lateral view, one half of the petiole pruned away by longitudinal section; G, scale from the abaxial face of the base of the petiole. A, Janssen et al. 2883 (P); B-G, Janssen et al. 2907 (P). Scale bars: A, 1 cm; B, G, 0.5 cm; C, 1 m; D, F, 5 cm; E, 0.1 cm.
FIG. 48 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 48. — Photographic documentation of field characters in Cyathea: A, B, D-F, habit; A', B', D'-F', trunk apex; A", B", D"-F", trunk surface at breast height. A-A", Cyathea meridionalis Janssen & Rakotondr.; B, B', C. ligulata Baker; C, trunk apex of C. acutula Tardieu var. acutula; D-D", C. acutula Tardieu var. deltoidea Janssen & Rakotondr.; E-E", C. appendiculata Baker; F-F", C. costularis Bonap. A, A", E, F, uncollected; A', Janssen et al. 2599; B, B', Janssen et al. 2494; C, Janssen et al. 2401; D-D", Janssen et al. 2788; E', E", Janssen et al. 2764; F', Janssen et al. 2489; F", Janssen et al. 2518. Vouchers at P. Photos B, C, F, A', B', F', F" by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A, B, D-F, 1 m; C, A', B', D'-F', A", D"-F", 10 cm.
FIG. 49 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 49. — Photographic documentation of field characters in Cyathea: A-E, habit; A'-E', trunk apex; A"-E", trunk surface at breast height. A-A", Cyathea hebes Janssen & Rakotondr.; B-B", C. serratifolia Baker; C-C", C. tsaratananensis C.Chr.; D-D", C. viguieri Tardieu; E-E", C. approximata Bonap.; E', left: apex of a form with distant petioles (apical view), right: apex of a form with close standing petioles (lateral view). A-A", Janssen et al. 2519; B, B", C, C', E-E", uncollected; B', Janssen et al. 2538; C", Janssen et al. 2446; D-D", Janssen et al. 2575. Vouchers at P. Photos A-A", B-B", C-C", E, E' right, E" by G. Rouhan (MNHN); all other photos by T. Janssen. Scale bars: A-E, 1 m; A'-E', A"-B", D", E", 10 cm; C", 5 cm.
FIG. 19 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 19. — Cyathea viguieri Tardieu: A, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view; B, scale from the abaxial face of the base of the petiole; C, habit; D, pinnae abaxially with a fragment of the rachis, sori only partly indicated; E, scale from the lateral face of the middle part of the petiole; F, pinnules abaxially with a fragment of the costa; G, sori with dark brown, coriaceous indusia; H, leaf scars and trunk surface. A, B, D-G, Janssen et al. 2575 (P); C, H, Janssen et al. 2380 (P). Scale bars: A, D, 5 cm; B, E, 0.5 cm; C, 1 m; F, H, 1 cm; G, 0.1 cm.
FIG. 18. — Cyathea tsaratananensis C in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 18. — Cyathea tsaratananensis C.Chr.: A, pinnules abaxially with a fragment of the costa, note that the sterile pinnule is more strongly crenate; B, leaf scars and trunk surface, form with close standing petioles; C, habit; D, leaf scars and trunk surface, common form; E, segment from the basal part of the petiole with scales; F, scales from the base of the petiole (left: strongly indurated, pyramidal scale; right: less strongly indurated scale); G, pinna abaxially with a fragment of the rachis, fertile in its lower half only. A, G, Rakotondrainibe et al. 5181 (P); B, Janssen et al. 2456 (P); C-F, Janssen et al. 2446 (P). Scale bars: A, B, D, E, 1 cm; C, 1 m; F, 0.1 cm; G, 5 cm.
FIG. 20 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 20. — Cyathea approximata Bonap.: A, basal part of the leaf (from the petiole base up to the first four pinna pairs), dorso-lateral view, note the minute scales on the petiole, the pinnae are pruned away on one side; B, pinnules abaxially with a fragment of the costa; C, habit; D, sori, note the dense scaly paraphyses inserted on the apex of the receptacle; E, pinnae abaxially with a fragment of the rachis, sori only partly indicated; F, scales from the base of the petiole (left: minute scale from the adult petiole; right: filiform scale from a crozier); G, leaf scars, note that the trunk surface is smooth. A, B, D-G, Janssen et al. 2449 (P); C, uncollected, photograph at P. Scale bars: A, 5 cm; B, E, G, 1 cm; C, 1 m; D, F, 0.1 cm.
FIG. 11 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 11. — Cyathea meridionalis Janssen & Rakotondr.: A, pinnules abaxially with a fragment of the costa, sori omitted from one pinnule; B, habit; C, pinnae abaxially with a fragment of the rachis, sori only partly indicated; D, basal part of the leaf (from the petiole base up to the first several pinna pairs), dorsal view; E, proximal part of a pinnule abaxially, with a dense indument of multicellular hairs; F, scale from the base of the petiole (left: dorsal view; right: lateral view, schematic); G, leaf scars and trunk surface. A-G, Janssen et al. 2603 (P). Scale bars: A, 0.5 cm; B, 1 m; C, D, 5 cm; E, F, 0.1 cm; G, 1 cm.
FIG. 17 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 17. — Cyathea serratifolia Baker: A, scale from the base of the petiole; B, habit (left: old plant, short and close standing petioles; right: young plant, long and distant petioles); C, leaf scars and trunk surface; D, pinnules abaxially with a fragment of the costa; E, pinna abaxially with a fragment of the rachis, sori only partly indicated. A, D, E, Janssen et al. 2537 (P); B, uncollected, photographs at P; C, Janssen et al. 2538 (P). Scale bars: A, 0.1 cm; B, 1 m; C, E, 5 cm; D, 1 cm.
FIG. 14 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 14. — Cyathea basirotundata Rakotondr. & Janssen: A, basal part of the leaf (from the petiole base up to the third pinna pair), lateral view, scales caducous; B, fertile pinnules abaxially with a fragment of the costa; C, sterile pinnules abaxially with a fragment of the costa; D, leaf scar and scaly trunk surface; E, fertile pinna with a fragment of the rachis; F, sterile pinna with a fragment of the rachis; G, sori in the upper half of a pinnule; H, scale from the base of the petiole. A-H, Rakotondrainibe 6377 (P). Scale bars: A, E, F, 5 cm; B-D, 1 cm; G, H, 0.1 cm.
FIG. 4. — A-E in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 4. — A-E, Cyathea marattioides Willd. ex Kaulf.; A, pinna abaxially with a fragment of the rachis, sori only partly indicated; B, habit; C, pinnules abaxially with a fragment of the costa; D, scale from the base of the petiole; E, trunk segment with leaf scars; F, G, C. tsilotsilensis Tardieu; F, pinnules abaxially with a fragment of the costa; G, scale from the base of the petiole; H-K, C. perrieriana C.Chr.; H, pinnules abaxially with a fragment of the costa; I, trunk segment with leaf scars; J, scale from the base of the petiole; K, habit. A-E, Janssen et al. 2488 (P); F, G, van der Werff 12905 (P); H-J, Janssen et al. 2503 (P); K, uncollected, photograph at P. Scale bars: A, 5 cm; B, K, 1 m; C, E, F, H, I, 1 cm; D, G, J, 0.5 cm.
FIG. 10 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 10. — Cyathea lisiae Janssen & Rakotondr.: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, pinnules abaxially with a fragment of the costa; C, proximal part of a pinnule abaxially with a dense indument of multicellular hairs and scale-topped hairs, note the rudimentary collar-like indusium; D, leaf scar; E, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view; F, bicolorous scale from the upper third of the petiole; G, concolorous scale from the base of the petiole. A-G, Rabarimanarivo 204 (P). Scale bars: A, E, 5 cm; B, 0.5 cm; C, F, G, 0.1 cm; D, 1 cm.
FIG. 9 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles
FIG. 9. — Cyathea ligulata Baker: A, pinnae adaxially with a fragment of the rachis, note that the auricles of the pinnules overlap the rachis; B, two strongly indurated pyramidal scales from the base of the petiole (left) and one narrowly triangular scale from the abaxial face of the costa (right); C, habit; D, basal part of a pinnule abaxially (at the junction of the costula with the costa) showing indusia as well as the densely scaly costa and costula; E, basal part of the leaf (from the petiole base up to the second pinna pair), dorsal view, basalmost pinnae are strongly reflexed; F, pinnules abaxially with a fragment of the costa, form with long pinnules with a crenate margin and an obtuse apex; G, pinnules abaxially with a fragment of the costa, form with short pinnules with a subentire margin and a rounded apex; H, leaf scars and spiny trunk surface. A, D, G, Rakotondrainibe et al. 5110 (P); B, E, Rakotondrainibe et al. 2057 (P); C, H, Janssen et al. 2494 (P); F, Rakotondrainibe et al. 2057 (P). Scale bars: A, E, 5 cm; B, F, G, 0.5 cm; C, 1 m; D, 0.1 cm; H, 1 cm.
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