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FIGURE 4 in New Eclipidrilus species (Annelida, Clitellata, Lumbriculidae) from southeastern North America

FIGURE 4. Eclipidrilus microthecus (G, specimen from Pettiford Creek, whole mount; all others from tributary to Naked Creek, A from a whole mount, others from sections). A–C, ectal part of spermathecal duct and vestibule: B shows the outer vestibule; C shows the ental part of the vestibule, with muscular ring. D, spermatheca. E, atrial duct and base of ampulla. F, sagittal section of atrial ampulla, with ental end and "prostate" to the right. G, atrial ampulla as seen in whole mount. H, transverse section of atrial ampulla, showing muscle layers and narrow lumen. I, ectal end of atrial duct and penial structure with (faint) penis. Scale bars: 100 μm.

FIGURE 5 in New Eclipidrilus species (Annelida, Clitellata, Lumbriculidae) from southeastern North America

FIGURE 5. Eclipidrilus macphersonae (A, E, specimens from Lumber River; D, F, G, specimens from Naked Creek; B, C, specimen from Flat Creek). A–B, lateral view of anterior segments. C, reproductive segments IX–XI, from a mature worm, facing right. D, spermatheca of a recently-mated worm, with sperm only in the ectal part. E, reproductive segments IX–X, showing genital pores and spermatheca of a fully mature worm, facing right. F, reproductive segments IX–XI, from an unmated worm, facing left. G, paired chaetae: from left to right, segment VI (ventral pair), XII (dorsal), and XX (ventral). Scale bars: A–B 500 μm, C–F 200 μm, G 50 μm.

FIGURE 3 in New Eclipidrilus species (Annelida, Clitellata, Lumbriculidae) from southeastern North America

FIGURE 3. Eclipidrilus microthecus (A–B, C, F, specimens from tributary to Naked Creek; D–E, specimens from Flat Creek). A–B, lateral view of anterior segments. C, paired chaetae: from left to right, segment IV (dorsal pair) and V (ventral). D, paired chaetae: IV (ventral) and XII (dorsal). E–F, reproductive segments IX–XI, from dissected worms; both worms facing right. G, ectal end of atrial duct and penis, from a whole mount. Scale bars: A–B 400 μm, C–D 50 μm, E–F 200 μm.

FIGURE 2 in New Eclipidrilus species (Annelida, Clitellata, Lumbriculidae) from southeastern North America

FIGURE 2. Eclipidrilus breviatriatus (all sagittal sections, specimens from tributary to Naked Creek). A–B, spermathecae. C, spermathecal ampulla. D, spermathecal duct and pore. E, entire atrium. F, transverse-diagonal section of atrial ampulla. G–J, sagittal sections of atrial ampullae; lumen expanded in H, large (glandular?) cells next to atrium in I–J. K–L, sagittal sections of atrial ducts and penes. Scale bars 100 μm.

FIGURE 8. R. diespluviae n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 8. R. diespluviae n. sp. from Rainy Day Creek, Idaho (A, C–D), from Cold Spring Creek, Idaho (B), from Hurst Creek, Washington (E–F), and from Orogrande Creek, Idaho (G–H), reproductive structures. A, F, G. Spermatheca, sagittal view. B–E, H. Penial bulb and ectal end of atrium in X, sagittal view from medial side; penial bulb (E) is everted. Scale bars 300 Μm (all drawings to same scale).

FIGURE 7. R. diespluviae n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 7. R. diespluviae n. sp. from Rainy Day Creek, Idaho, reproductive structures. A. Atrium and vasa deferentia, sagittal view from a dissected worm (X–XVII, facing left). B. Spermathecae in VIII, reconstructed from transverse sections (arrows indicate positions of chaetae). C. Penial structures in X, reconstructed from transverse sections. Scale bar (B–C) 400 Μm.

FIGURE 9. R. diespluviae n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 9. R. diespluviae n. sp. from Rainy Day Creek, reproductive structures; sagittal sections (A–B), transverse sections (C–I). A. Spermatheca. B. Detail of spermathecal bulb with very short duct at top. C. Spermathecal diverticulum. D. Spermathecal ampulla. E. Penial sac and base of penis, containing end of atrial duct. F. Penial sac and tip of penis. G–I. Atrium and vasa deferentia: near the ectal end, in XI (G); near the transition to the prostate-covered section (H); and in the ental, prostate-covered section, with 1 vas deferens embedded in the atrial muscle (I). Scale bars (A, C) 200 Μm, (B, D–I) 100 Μm.

FIGURE 5. R. klamathensis n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 5. R. klamathensis n. sp. from Upper Klamath Lake, reproductive organs; sagittal sections (A–D, F), transverse sections (E, G–I). A. Spermathecal bulb, duct, and ampulla joining the gut. B. Spermathecal bulb and duct, with part of a diverticulum. C. Penial bulb and sac, with a retracted penis. D–E. Partially retracted penes, within penial bulbs. F. Atrium, vas deferens and posterior male funnel at 10/11. G. Ectal part of atrium in X, with vasa deferentia free in sperm sac. H. Ental part of atrium with vasa deferentia free within prostate layer; the small, anterior vas deferens approaching the atrium. I. Atrium near the ental end; the posterior vas deferens is approaching the atrium. Scale bars (A–F) 200 Μm, (G–I) 100 Μm.

FIGURE 6. R. diespluviae n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 6. R. diespluviae n. sp., external characters; worms from Rainy Day Creek, Idaho (A, B, E), from Hurst Creek, Washington (D), and from North Fork Clearwater River, Idaho (C). A–C. Ventral views of anterior ends, showing positions of genital pores. D. Lateral view of a worm with everted penes (proboscis was broken off). E. Chaetae, from left to right: a dorsal pair from XI, a ventral pair from XI, and a pair from about the posterior 1/3 of the worm. Scale bars (A–D) 2 mm, (E) 100 Μm.

FIGURE 4. R. klamathensis n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 4. R. klamathensis n. sp., reproductive organs drawn from dissected worms; from Upper Klamath Lake (A–F), from Mokelumne River (G–I). A–C, G–H, spermathecae; D–F, ectal ends of male ducts with penial bulbs; I, atrium and penial bulb. A. Spermatheca with a slightly branched diverticulum. B. An unusual specimen with highly branched spermathecal diverticula. C. A typical specimen with unbranched spermathecal diverticula. D. A completely retracted penial bulb. E. A slightly everted penial bulb. F. An everted penial bulb (same specimen as Fig. 2C). G. Mature spermatheca. H. Spermatheca from partiallymature worm. I. A relatively straight atrium with retracted penis (anterior vas deferens not shown). Scale bar 300 Μm (all drawings to same scale).

FIGURE 3. R. klamathensis n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 3. R. klamathensis n. sp., reproductive organs, from Upper Klamath Lake (A, C–E) and from Mokelumne River (B). A–B. Segments VIII–XI, from longitudinally-dissected specimens (facing left). C. Atrium and male ducts from a partiallymature worm (facing left). D. Spermathecae in VIII, reconstructed from transverse sections. E. Penial structures in X, reconstructed from transverse sections. Scale bars 500 Μm (all drawings to same scale). Arrows: position of chaetae (not in sectional plane here).

FIGURE 2. R. klamathensis n in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 2. R. klamathensis n. sp. from Upper Klamath Lake (A–F, I–J) and from Mokelumne River (G–H). A–D. Lateral and ventral views of anterior segments of mature worms; penes everted in C–D. E–F. Lateral and ventral views of a specimen with partially-developed reproductive pores. G–H. Lateral views of a mature and immature worm. I. Lateral blood vessels in XXV, worm facing right; at lower right, the broken end of the anterior vessel (small arrow) indicates the junction with the perivisceral sinus. J. Chaetae: from left to right, a pair from mid-body region, a ventral pair from segment VIII, and a pair from about the posterior 1/4 of a worm. Scale bars (A–H) 2 mm, (I) 400 Μm, (J) 200 Μm.

FIGURE 1. R in Rhynchelmis subgenus Sutroa Eisen new rank, with two new species from western North America (Annelida, Clitellata, Lumbriculidae)

FIGURE 1. R. rostrata from the type locality, Mountain Lake, San Francisco (A–B) and from Sycan Marsh, Klamath watershed (C). A. spermatheca. B. Ectal end of male duct, showing the conical penis retracted within bulb, ectal part of the atrium with thick muscle layer, and the vasa deferentia. C. Anterior end of a whole worm, with everted penes. Scale bars (A–B) 400 Μm, (C) 2 mm.

FIGURE 1 in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 1. Sampling sites in Tibet. A. Map of China. B. Sampling sites of Yarlung Zangbo River, Lhasa River, Niyang River, Lake Yamzho Yumco, Lake Namco and Lalu Wetland, respectively.

FIGURE 4. Nais longidentata n in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 4. Nais longidentata n. sp. A–C. Ventral chaetae (III, VI, posterior, respectively). D. Needles. E. Penial chaetae. F. Lateral view of sexual organs in segments V–VI. Scale bar: A–D 20 Μm; E 50 µm, F 100 µm.

FIGURE 2 in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 2. Chaetogaster diaphanus (Gruithuisen, 1828). A–C. Ventral chaetae (A: VI; B: II; C: posterior). D–F. Penial chaetae. G. Ventral view of sexual organs in segments IV–VI. Scale bars: A–F 40 µm, G 160 µm.

FIGURE 6 in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 6. Rhyacodrilus stephensoni Černosvitov, 1942. A. Spermathecal chaeta. B. Dorsal chaeta. C. Lateral view of male duct and spermatheca in segments X–XI. Scale bars: A–B 20 Μm, C 80 Μm.

FIGURE 3 in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 3. Nais elinguis Müller, 1774. A. Needle chaeta. B. Penial chaeta. C. Ventral view of sexual organs in segments V– VI. Scale bars: A–B 20 µm, C 40 µm.

FIGURE 5 in Records of Naididae and Lumbriculidae (Clitellata) from Tibet, China, with description of a new species of Nais

FIGURE 5. SEM (A–E) and optical microscope (F) micrographs of Nais longidentata n. sp. A–B. Ventral chaetae in IX (A) and XIII (B). C. Hair in VII. D–E. Needles in X (D) and a posterior segment (E). F. Lateral view of anterior body. Scale bars: A,B, D 20µm, C 5µm, E 10 µm.

FIGURE 4. Uktena riparia n in Uktena riparia n. gen., n. sp. (Annelida, Clitellata, Lumbriculidae), a new spermatophore-producing oligochaete

FIGURE 4. Uktena riparia n. sp. from Flat Creek (the type locality), from sagittal sections (A–E and J–L), or dissections (F– I). A. Ectal part of spermathecal ampulla, with sperm lined up along epithelium. B. Ental part of spermathecal ampulla with weakly-stained (old?) sperm. C. Copulatory organ and gland in spermathecal bursa. D. Muscle layer of spermathecal bursa wall. E. Small glands and muscle layer forming the wall of male bursa. F. Gland opening on papilla within male bursa. G. Spermatophore forming in atrial duct, with darkly-staining sperm (composite photo). H. Atrial ampulla and duct with coiled sperm bundles inside. I. Spermatophore protruding from male pore. J–L. Atrial ampullae; (K) showing detail of atrial wall, with prostate glands penetrating muscle layer (arrows); (L) showing elongate prostate glands with secretions.